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26 April 2001 Checklist of Species of the Subfamily Masarinae (Hymenoptera: Vespidae)
JAMES M. CARPENTER
Author Affiliations +
Abstract

A checklist of species in the pollen wasp subfamily Masarinae is presented, including synonyms and distributional summaries. The 297 valid species and 15 additional subspecies are arranged in 14 genera, divided into two tribes, Gayellini and Masarini, with the latter divided into the subtribes Priscomasarina, Paragiina, and Masarina. Thirty-two new combinations are made.

INTRODUCTION

The most recent monograph of the Masarinae is by Richards (1962). Since that work there have been numerous changes to the classification of these wasps. The family group arrangement is now different: Richards treated Masarinae in the present sense as a family, with three subfamilies: Euparagiinae, Gayellinae, and Masarinae. Masarinae was divided into tribes, Paragiini and Masarini. Carpenter (1981) reduced Masaridae to a subfamily of Vespidae, removing Euparagiinae, which is the sister-group to all other extant vespids, reducing Richards' Gayellinae and Masarinae to tribes, and sinking Richards' Paragiini and Masarini. Carpenter (1997) reconstituted these latter groups and recognized them as subtribes within Masarini, to which F. W Gess (1998) added another subtribe, Priscomasarina. The generic classification has also undergone considerable change: Giordani Soika (1974), Snelling (1986), and F. W. Gess (1998) each added a genus to the 18 recognized by Richards, while Carpenter (1989), van der Vecht and Carpenter (1990), and Carpenter (1997) synonymized eight. And of course, there have been numerous species descriptions and synonymies published since Richards (1962).

More recently, S. K. Gess (1996) reviewed the Masarinae and provided a list of described species, along with distributional summaries. That list is useful; however, it lacks citations of the original references, limiting its taxonomic value. There have been a number of descriptions and synonymies since that list was published; moreover, preparation of the present checklist has revealed a few errors in both Richards (1962) and S. K. Gess (1996).

The present checklist follows the generic classification in van der Vecht and Carpenter (1990), with one amendment. The generic synonymies in van der Vecht and Carpenter's catalog were based on my ongoing phylogenetic studies in the Vespidae. Only parts of the cladistic study of Masarinae have as yet been published (Carpenter, 1989, 1997; see summary in Carpenter, 1993). Because the rationale for most of the synonymy was not yet published, S. K. Gess (1996) did not follow them in her list. Carpenter (1997) subsequently detailed the basis for generic reclassification in the Paragiina. I will summarize the rationale for the remaining synonymy here.

The monophyly of Trimeria + Microtrimeria is shown by the forewing cu-a distad of the fork of M and CuA by a distance nearly equal to its length, a single occipital carina, the palpal formula 2:3, loss of the lateral pronotal groove, the short pyriform tegula, the posteriorly lamellate scutellum, the female foretarsi with lobes of unique form, the hooked hairs on the female foretarsi, and the compressed mid- and hindcoxae. Both genera are evidently monophyletic, as is the Trimeria subgenus Willinkia Richards, but that does not mean that their recognition is warranted. The characters by which all these taxa differ are for the most part variable within other genera, and so their recognition contributes little to the process of efficient diagnosis (Farris, 1979). Accordingly, they are treated as synonyms.

Richards (1962) recognized two subgenera within Celonites, but whereas Eucelonites is defined by the autapomorphy of an axillary process, the typical subgenus is defined only by its absence. It is evidently paraphyletic, and Eucelonites is thus synonymized with Celonites.

The monophyly of Quartinia + Quartinioides + Quartiniella is shown by the forewing marginal cell appendiculate with a nebulous vein, the forewing cu-a oblique, the hindwing preaxillary excision deep, the single long occipital carina, and the palpal formula 1:3. Both Quartiniella and Quartinioides are primarily based on partitioning a trend in the reduction of wing venation. Quartiniella is defined by the loss of forewing 3rs-m and 2m-cu; Quartinioides has 2m-cu present but attenuate and interrupted, whereas Quartinia has it complete. Other features listed in Richards' (1962: 127) key mostly vary within these taxa or are erroneous (e. g., Quartinioides has two hindtibial spurs). The remainder are consistent with the inference that Quartinia is paraphyletic in terms of Quartinioides, which is in turn paraphyletic in terms of Quartiniella (e. g., Quartinioides is also separated from Quartinia by the preaxillary excision deep, the jugal lobe reduced, and the stipes medially produced, all characters shared with Quartiniella). In any case, the formal taxonomic partitioning of essentially continuous variation should be rejected, and Quartinioides and Quartiniella are accordingly synonymized with Quartinia.

Finally, I previously (van der Vecht and Carpenter, 1990) treated Masarina as a synonym of Jugurtia. Both genera have autapomorphies, but considering the few species of Masarina recognized at that time (i. e., four; see Richards, 1962, Gess and Gess, 1988), I did not regard the features by which the genera differ as significant as those which they share. Further, the recognition of a genus Masarina might be confusing because that name is also used for a subtribe. F. W. Gess (1997) has since described six more species of Masarina and pointed out additional diagnostic features, and I agree that maintaining the genera is now justified. A total of 14 genera (and no subgenera) for the Masarinae is thus recognized here.

The arrangement of the checklist is alphabetical. Each species and subspecies is followed by the original citation and then by synonyms, which are listed chronologically. The status given each taxon generally follows the most recent author to treat that taxon. I do not regard the recognition of subspecies as taxonomically useful, but as such taxa are a mixture of synonyms and valid phylogenetic species, each case should be studied before appropriate corrective action is taken. Misspellings are indicated by exclamation marks in brackets. To facilitate ready reference, each publication is cited in abbreviated form. Where two dates are listed, the first is the true date of publication, so far as can be determined, with the date following in parentheses being the date printed on the publication. The sex of the type specimens, if stated in the description, is then listed. The original locality is quoted for each description, and the type depository is then listed in parentheses. If the sex of the holotype or lectotype is known, this is listed with the depository, except where a single sex was specified in the original description, in which case this information is not repeated, or where syntypes have not had a lectotype designated (a few exceptions to these rules are corrections to original descriptions). A code word is used to designate most type depositories in the checklist, typically the name of city in which the collection is located; this precedes the name of the museum or institution. Museums or institutions where type specimens are kept are as follows.

  • Berlin: Museum für Naturkunde der Humboldt-Universität, Berlin, Germany

  • BOL: Biologiezentrum des Oberösterreichischen Landesmuseums Linz, Austria

  • Bremen: Übersee-Museum, Bremen, Germany

  • Budapest: Hungarian Natural History Museum, Budapest, Hungary

  • Buenos Aires: Museo Argentino de Ciencias Naturales “Bernardino Rivadavia,” Buenos Aires, Argentina

  • Cambridge: Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA

  • Canberra: Australian National Insect Collection, CSIRO, Canberra, Australia

  • Cape Town: South African Museum, Cape Town, South Africa

  • Copenhagen: Zoologisk Museum, Kobenhavns Universitet, Copenhagen, Denmark

  • Davis: Bohart Museum of Entomology, University of California, Davis, California, USA

  • Geneva: Muséum d'Histoire Naturelle, Geneva, Switzerland

  • Genova: Museo Civico di Storia Naturale “Giacomo Doria,” Genoa, Italy

  • Grahamstown: Albany Museum, Grahamstown, South Africa

  • Hamburg: Zoologisches Museum, Universität Hamburg, Hamburg, Germany

  • Helsinki: Finnish Museum of Natural History, Helsinki, Finland

  • IPC: Instituto Pedagógico de Chile [Universidad de Chile, Santiago, Chile]

  • Leiden: Nationaal Natuurhistorisch Museum, Leiden, the Netherlands

  • Linz: coll Gusenleitner, Pfitznerstrasse 31, A-4020 Linz/Donau, Austria

  • London: Natural History Museum, London, England

  • Madrid: Museo Nacional de Ciencias Naturales, Madrid, Spain

  • Moscow: Zoological Museum, Moscow State University, Moscow, Russia

  • New York: American Museum of Natural History, New York, New York, USA

  • Oxford: Hope Entomological Collections, Oxford University, Oxford, England

  • Paris: Muséum National d'Histoire Naturelle, Paris, France

  • Perth: Western Australian Museum, Perth, Australia

  • Philadelphia: Academy of Natural Sciences, Philadelphia, Pennsylvania, USA

  • Pretoria: Transvaal Museum, Pretoria, South Africa

  • St. Petersburg: Zoological Institute, Russian Academy of Sciences, St Petersburg, Russia

  • San Francisco: California Academy of Sciences, San Francisco, California, USA

  • San Lorenzo: Museo Nacional de Historia Natural del Paraguay, San Lorenzo, Paraguay

  • Santiago: Museo Nacional de Historia Natural, Santiago, Chile

  • Sofia: National Museum of Natural History, Bulgarian Academy of Sciences, Sofia, Bulgaria

  • Tucumán: Fundación y Instituto Miguel Lillo, Tucumán, Argentina

  • Turin: Museo di Zoologia Sistematica dell'Università di Torino, Turin, Italy

  • Uppsala: Zoological Museum, Uppsala University, Uppsala, Sweden

  • Venice: Museo Civico di Storia Naturale, Venice, Italy

  • Vienna: Naturhistorisches Museum Wien, Vienna, Austria

  • Washington: National Museum of Natural History, Smithsonian Institution, Washington, DC, USA

  • Zürich: Eidgenössiche Technische Hochschule, Zürich, Switzerland

A few smaller private collections are not included in the above list, being mostly collections whose present whereabouts are doubtful.

Distributional data are summarized by country in the final paragraph for each species. An asterisk denotes a new locality record.

SUBFAMILY MASARINAE LATREILLE, 1802

  • Masarides Latreille, 1802, Hist. Nat. Crust. Insect. 3: 367, family. Type genus: Masaris Fabricius, 1793.

  • Mazaridinae Schulz, 1911, Zool. Ann. Würzburg 4: 199. Unnecessary emendation of Masarinae Latreille.

  • Celonitinae Börner, 1919, Biol. Zentralbl. 39(4): 186. Type genus: Celonites Latreille, 1802.

  • Saryarinae Kuznetzov, 1923, J. Turkestan Branch Russ. Geogr. Soc. 16: 127. Type genus: Saryara Kuznetzov, 1923.

  • Tribe Gayellini Bradley, 1922

  • Gayellinae Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 374, 376, 380 (key), subfamily. Type genus: Gayella Spinola, 1851.

  • Genus Gayella Spinola

  • Gayella Spinola, 1851, in Gay, Hist. Fis. Polit. Chile, 6: 328, genus. Type species: Gayella eumenoides Spinola, 1851, by monotypy.

  • Gazella[!]Reed, 1894, Proc. Zool. Soc. London 1893: 685.

  • araucana Willink

  • Gayella araucana Willink, 1956, Bol. Acad. Nac. Cienc. 39: 341, 342, 346, figs. 3, 4, female, male –“Chile, Prov. Santiago: Renca” (holotype female, IPC).

  • Distribution: Chile: Atacama, Coquimbo, Valparaíso, Santiago, O'Higgins, Maule, Biobío.

  • eumenoides Spinola

  • Gayella eumenoides Spinola, 1851, in Gay, Hist. Fis. Polit. Chile. Zool. 6: 333, pl. II, fig. 2, male, female –“provincias del norte y sobretodo en Santa Rosa” (lectotype female, Paris).

  • Gayella sicheliana von Schulthess, 1910, Deutsch. Entomol. Z.: 189, as a synonym of eumenoides, attributed to de Saussure.

  • Distribution: Chile: Atacama, Coquimbo, Valparaíso, Santiago, O'Higgins, Maule.

  • luispenai Willink and Ajmat de Toledo

  • Gayella luispenai Willink and Ajmat de Toledo, 1979, Acta Zool. Lilloana 35: 427, 428, 429, figs. 1, 6, 7, male, female –“Riconada, Jujuy, Argentina” (holotype male, Tucumán).

  • Distribution: Argentina: Jujuy; Bolivia: Potosí.

  • mutilloides de Saussure

  • Gayella mutilloides de Saussure, 1855, Ét. Fam. Vesp. 3: 114, female –“Le Chili” (London).

  • Gayella odyneroides; von Schulthess, 1910, Deutsch. Entomol. Z.: 189. Lapsus calami.

  • Gayella mutilloides nigerrima Giordani Soika, 1960 (1958), Boll. Mus. Civ. Stor. Nat. Venezia 11: 80, female –“Cile” (Venice).

  • Distribution: Chile: Valparaíso, Santiago, Biobío, Araucanía; Argentina: Chubut, Neuquén, Rio Negro.

  • patagonica Willink

  • Gayella patagonica Willink, 1956, Bol. Acad. Nac. Cienc. 39: 341, 342, 350, figs. 9–11, female, male –“Esquel, Chubut, Argentina” (holotype female, Tucumán).

  • Gayella cerceroides Giordani Soika, 1960 (1958), Boll. Mus. Civ. Stor. Nat. Venezia 11: 82, female –“N.W. Patagonia, 1000–3000 piedi” (London).

  • Distribution: Chile: Santiago; Argentina: Chubut, Mendoza, Rio Negro.

  • reedi Willink

  • Gayella reedi Willink, 1963, Acta Zool. Lilloana 19: 385, 1 fig., female –“Casa Blanca, Valparaíso, Chile” (San Francisco).

  • Distribution: Chile: Atacama, Coquimbo, Valparaíso.

  • Genus Paramasaris Cameron

  • Paramasaris Cameron, 1901, Trans. Am. Entomol. Soc. 27: 311, genus. Type species: Paramasaris fuscipennis Cameron, 1901, by monotypy.

  • Zethoides Cameron, 1904, Trans. Am. Entomol. Soc. 30: 93, genus, non Zethoides Fox, 1899. Type species: Zethoides flavolineatus Cameron, 1904, by monotypy.

  • Plesiozethus Cameron, 1905, Entomologist 38: 269, replacement name for Zethoides Cameron.

  • Metazethoides Schulz, 1906, Spolia Hym.: 213, unnecessary replacement name for Zethoides Cameron.

  • Paragayella Giordani Soika, 1974, Boll. Mus. Civ. Stor. Nat. Venezia 25: 87, 89, 99, genus. Type species: Paragayella richardsi Giordani Soika, 1974, by original designation and monotypy.

  • brasiliensis Giordani Soika

  • Paramasaris brasiliensis Giordani Soika, 1974, Boll. Mus. Civ. Stor. Nat. Venezia 25: 105, female, male –“Brasile: Nova Teutonia, Santa Catarina” (holotype female, Cambridge).

  • Distribution: Brazil: Santa Catarina; Argentina: Misiones.

  • cupreus Giordani Soika

  • Paramasaris cupreus Giordani Soika, 1974, Boll. Mus. Civ. Stor. Nat. Venezia 25: 104, 106, female –“Columbia: Caqueta, Florencia, 480 m.” (London).

  • Distribution: Colombia: Caqueta, Magdalena, Putumayo; Peru: Loreto.

  • fuscipennis Cameron

  • Paramasaris fuscipennis Cameron, 1901: 312, female –“Santa Fé Mountains, New Mexico” (London).

  • Zethoides flavolineatus Cameron, 1904, Trans. Am. Entomol. Soc. 30: 94, male –“Panama (Pacific side)” (London).

  • Distribution: Colombia; Panama; Guatemala; Mexico: Jalisco, Sinaloa.

  • richardsi (Giordani Soika)

  • Paragayella richardsi Giordani Soika, 1974, Boll. Mus. Civ. Stor. Nat. Venezia 25: 88, fig. 2, pl. II, 101, female –“Brasile: Mato Grosso, Serra Roncados, R. S. Base Camp” (London).

  • Distribution: Brazil: Goiás, Mato Grosso.

  • Tribe Masarini Latreille, 1802

    Subtribe Priscomasarina Gess, 1998

  • Priscomasarina Gess, 1998, J. Hym. Res. 7(2): 296, 299, subtribe. Type genus: Priscomasaris Gess, 1998.

  • Genus Priscomasaris Gess

  • Priscomasaris Gess, 1998, J. Hym. Res. 7(2): 296, genus. Type species: Priscomasaris namibiensis Gess, 1998, by original designation and monotypy.

  • namibiensis Gess

  • Priscomasaris namibiensis Gess, 1998, J. Hym. Res. 7(2): 296, 300, figs. 2–9, female, male –“Namibia: between Palm and Khorixas (20.17S 14.05 E)” (holotype female, Grahamstown); also from five other localities in Namibia.

  • Distribution: Namibia.

  • Subtribe Paragiina Bradley, 1922

  • Paragiini Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 387 (key), tribe. Type genus: Paragia Shuckard, 1837.

  • Genus Metaparagia Meade-Waldo

  • Metaparagia Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 748, genus. Type species: Paragia pictifrons Smith, 1857, by original designation.

  • Riekia Richards, 1962, Revision. Study Masarid Wasps: 4, 53, 54, genus. Type species: Riekia nocatunga Richards, 1962, by original designation and monotypy.

  • Rolandia Richards, 1962, Revision. Study Masarid Wasps: 4, 53, 57, genus. Type species: Paragia maculata Meade-Waldo, 1910, by original designation and monotypy.

  • Ammoparagia Snelling, 1986, Contrib. Sci. 378: 1, 2, genus. Type species: Ammoparagia hua Snelling, 1986, by original designation and monotypy.

  • angulata (Richards)

  • Riekia angulata Richards, 1968, J. Aust. Entomol. Soc. 7: 101, female –“QUEENSLAND … Cunnamulla” (Canberra); also from New South Wales.

  • Distribution: Australia: Queensland, New South Wales.

  • borreriae (Snelling)

  • Rolandia borreriae Snelling, 1986, Contrib. Sci. 378: 1, 8–9, 11, figs. 11, 17, 24, male, female –“Cooper Creek (12°06;prS, 133°04;prE), 19 km E by S Mt. Borradaile, Northern Territory” (holotype male, Perth).

  • Distribution: Australia: Northern Terrritory.

  • confluens (Snelling)

  • Paragia confluens Snelling, 1986, Contrib. Sci. 378: 1, 14, fig. 28, female –“Western Australia … 36 km ESE Minnie Creek Homestead, 24°38;prS, 115°42;prE” (Perth).

  • Distribution: Australia: Western Australia.

  • doddi Meade-Waldo

  • Metaparagia doddi Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 748, female –“Cairns, N. Queensland” (London).

  • Distribution: Australia: Queensland.

  • houstoni (Snelling)

  • Rolandia houstoni Snelling, 1986, Contrib. Sci. 378: 1, 8, figs. 9, 12, 14, 15, 18, 22, 23, 26, 29, male, female –“Western Australia … 10 km ESE Meedo Homestead (25°40;prS, 114°37;prW)” (holotype male, Perth).

  • Distribution: Australia: Western Australia.

  • hua (Snelling)

  • Ammoparagia hua Snelling, 1986, Contrib. Sci. 378: 1, 3, 5, figs. 1–8, male, female –“Western Australia … 43 km ENE Landor Homestead (25°08;prS, 116°54;prE)” (holotype male, Perth).

  • Distribution: Australia: Western Australia.

  • maculata (Meade-Waldo)

  • Paragia maculata Meade-Waldo, 1910, Ann. Mag. Nat. Hist. (8) 5: 31, 32, female, male –“Western Australia” (lectotype female, London).

  • Distribution: Australia: Western Australia.

  • nocatunga (Richards)

  • Riekia nocatunga Richards, 1962, Revision. Study Masarid Wasps: 54, figs. 55–59, female –“New South Wales: Nocatunga (7 miles N. of)” (Canberra).

  • Distribution: Australia: New South Wales.

  • pictifrons (Smith)

  • Paragia pictifrons Smith, 1857, Cat. Hym. Brit. Mus.: 2, female –“Swan River” (London).

  • Distribution: Australia: Western Australia.

  • Genus Paragia Shuckard

  • Paragia Shuckard, 1837, Trans. Entomol. Soc. London 2: 81, genus. Type species: Paragia decipiens Shuckard, 1837, by monotypy.

  • Alpha Dalla Torre, 1904, Genera Insect. 19: 4, name for division I of genus Paragia Shuckard in de Saussure, 1855, Ét. Fam. Vesp. 3: 54 (3 species). Non Alpha de Saussure, 1854.

  • Beta Dalla Torre, 1904, Genera Insect. 19: 4, name for division II of genus Paragia Shuckard in de Saussure, 1855, Ét. Fam. Vesp. 3: 56 (5 species). Non Beta de Saussure, 1854.

  • Cygnaea Richards, 1962, Revision. Study Masarid Wasps: 4, 53, 60, subgenus of Paragia Shuckard. Type species: Paragia vespiformis Smith, 1865, by original designation and monotypy.

  • Paragiella Richards, 1962, Revision. Study Masarid Wasps: 4, 53, 67, subgenus of Paragia Shuckard. Type species: Paragia odyneroides Smith, 1850, by original designation.

  • australis australis de Saussure

  • Paragia australis de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xxi, sex not stated –“Nova Hollandia” (Paris).

  • Distribution: Australia: New South Wales.

  • australis borealis Richards

  • Paragia australis borealis Richards, 1962, Revision. Study Masarid Wasps: 67, 71, female, male –“Queensland: 17 miles west of Morven” (holotype female, Canberra).

  • Distribution: Australia: Queensland.

  • bicolor de Saussure

  • Paragia bicolor de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xxi, sex not stated –“Nova Hollandia” (Paris).

  • Distribution: Australia: New South Wales.

  • calida Smith

  • Paragia calida Smith, 1865, Trans. Entomol. Soc. London (3) 2: 392, male –“Adelaide” (London).

  • Distribution: Australia: South Australia, Western Australia.

  • deceptor Smith

  • Paragia deceptor Smith, 1862, Trans. Entomol. Soc. London (3) 1: 56, female –“Australia” (London).

  • Paragia deceptrix; Schulz, 1906, Spolia Hym.: 213. Unnecessary emendation.

  • Distribution: Australia: Queensland, New South Wales.

  • decipiens aliciae Richards

  • Paragia decipiens aliciae Richards, 1962, Revision. Study Masarid Wasps: 63, 64, female, male –“Northern Territories: Alice Springs” (holotype female, London).

  • Distribution: Australia: Northern Terrritory, Queensland.

  • decipiens decipiens Shuckard

  • Paragia decipiens Shuckard, 1837, Trans. Entomol. Soc. London 2: 82, pl. VIII, fig. 3, female –“New South Wales” (London).

  • Distribution: Australia: New South Wales, Victoria, South Australia.

  • generosa Richards

  • Paragia generosa Richards, 1962, Revision. Study Masarid Wasps: 67, 69, female –“Queensland: 30 miles S.S.W. of Ayr” (Canberra).

  • Distribution: Australia: Queensland.

  • hirsuta Meade-Waldo

  • Paragia hirsuta Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 749, male –“Cairns, N. Queensland” (London).

  • Distribution: Australia: Queensland.

  • magdalena Turner

  • Paragia magdalena Turner, 1908: 89, female –“MACKAY, Queensland” (London).

  • Distribution: Australia: Queensland.

  • mimetica Richards

  • Paragia mimetica Richards, 1968, J. Aust. Entomol. Soc. 7: 101, 102, male, female –“WESTERN AUSTRALIA … 10 m. W. Watheroo” (holotype male, Canberra).

  • Distribution: Australia: Western Australia.

  • monocesta Snelling

  • Paragia monocesta Snelling, 1986, Contrib. Sci. 378: 1, 15–16, figs. 27, 30, 31, female, male –“Western Australia … 10 km ESE Meedo Homestead, 25°40;prS, 114°37;prE” (holotype female, Perth).

  • Distribution: Australia: Western Australia.

  • morosa Smith

  • Paragia morosa Smith, 1868, Trans. Entomol. Soc. London 1868: 251, female –“Champion Bay” (London).

  • Distribution: Australia: Western Australia.

  • nasuta Smith

  • Paragia nasuta Smith, 1868, Trans. Entomol. Soc. London 1868: 252, female, male –“Champion Bay” (lectotype female, London).

  • Distribution: Australia: Queensland, Western Australia.

  • odyneroides Smith

  • Paragia odyneroides Smith, 1850, Trans. Entomol. Soc. London (2) 1: 41, pl. 5, fig. 2, male –“Hunter River, New Holland” (London).

  • Paragia bidens de Saussure, 1855, Ét. Fam. Vesp. 3: 59, male –“L'Adélaïde, Nouvelle –Hollande” (London).

  • Paragia praedator de Saussure, 1855, Ét. Fam. Vesp. 3: 59, female –“Le sud de la Nouvelle –Hollande” (London).

  • Distribution: Australia: Queensland, New South Wales, Victoria, South Australia.

  • oligomera Snelling

  • Paragia oligomera Snelling, 1986, Contrib. Sci. 378: 1, 13, female –“Western Australia … 10 km NE Wanneroo” (Perth).

  • Distribution: Australia: Western Australia.

  • perkinsi Meade-Waldo

  • Paragia perkinsi Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 750, female –“Cairns, N. Queensland” (London).

  • Distribution: Australia: Queensland.

  • propodealis Richards

  • Paragia propodealis Richards, 1968, J. Aust. Entomol. Soc. 7: 101, 104, female –“NEW SOUTH WALES … Caldwell” (Canberra).

  • Distribution: Australia: New South Wales.

  • schulthessi Turner

  • Paragia schulthessi Turner, 1936, Ann. Mag. Nat. Hist. (10) 18: 352, female –“Dedari, 40 miles west of Coolgardie, W. Australia” (London).

  • Distribution: Australia: Western Australia.

  • smithii de Saussure

  • Paragia tricolor Smith, 1850, Trans. Entomol. Soc. London (2) 1: 41, male, not female.

  • Paragia smithii de Saussure, 1854, Ét. Fam. Vesp. 3: pl. II fig. 1, female; 1855, Ét. Fam. Vesp. 3: 55 –“L'Australie Occidentale” (type depository unknown).

  • Distribution: Australia: New South Wales, Victoria, South Australia, Western Australia.

  • sobrina Smith

  • Paragia sobrina Smith, 1869, Trans. Entomol. Soc. London 1869: 309, female –“Champion Bay” (London).

  • Paragia excellens Smith, 1869, Trans. Entomol. Soc. London 1869: 309, female, male –“Champion Bay” (lectotype female, London).

  • Distribution: Australia: Western Australia.

  • tricolor Smith

  • Paragia tricolor Smith, 1850, Trans. Entomol. Soc. London (2) 1: 41, pl. 5, fig. 1, female not male –“Perth, W. Australia” (London).

  • Paragia saussurii Smith, 1857, Cat. Hym. Brit. Mus.: 2, female –“Perth” (London).

  • Paragia saussurei[!]; Meade-Waldo, 1910, Ann. Mag. Nat. Hist. (8) 5: 32; Richards, 1962, Revision. Study Masarid Wasps: 66, 294.

  • Distribution: Australia: Western Australia.

  • venusta Smith

  • Paragia venusta Smith, 1865, Trans. Entomol. Soc. London (3) 2: 393, female –“Swan River” (London).

  • Paragia concinna Smith, 1868, Trans. Entomol. Soc. London 1868: 251, female –“Champion Bay” (London).

  • Distribution: Australia: Western Australia.

  • vespiformis Smith

  • Paragia vespiformis Smith, 1865, Trans. Entomol. Soc. London (3) 2: 393, female –“Swan River” (London).

  • Distribution: Australia: Western Australia.

  • walkeri Meade-Waldo

  • Paragia walkeri Meade-Waldo, 1910, Ann. Mag. Nat. Hist. (8) 5: 31, 33, male –“Adelaide River, Northern Territory” (London).

  • Distribution: Australia: Northern Terrritory, Queensland.

  • Subtribe Masarina Latreille, 1802

    Genus Celonites Latreille

  • Celonites Latreille, 1802, Hist. Nat. Crust. Insect. 3: 368, genus. Type species: Masaris apiformis Fabricius, 1793 (= Vespa abbreviata Villers, 1789), by monotypy.

  • Chelonites[!] Leach, 1815, “Entomology” in Edinburgh Encyclop. IX(1): 154.

  • Coelonites[!] Blanchard, 1840, Hist. Nat. Anim. Artic. 3: 387.

  • Paracelonites Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 108, subgenus of Celonites Latreille (6 species). Unavailable; no type species designated.

  • Eucelonites Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 112, subgenus of Celonites Latreille (4 species). Unavailable; no Type species: designated.

  • Eucelonites Richards, 1962, Revision. Study Masarid Wasps: 214, 215, subgenus of Celonites Latreille. Validation by type selection of Eucelonites Kostylev, 1935. Type species: Celonites cyprius de Saussure, 1854, by original designation.

  • abbreviatus abbreviatus (Villers)

  • Vespa abbreviata Villers, 1789, C. Linn. Entomol. 3: 281, sex not stated –“Europa. In Gallia Australiori” (destroyed).

  • Chrysis dubia Rossi, 1790, Fauna Etrusca 2: 77, pl. VII, figs. x–xi, sex not stated [female] –“Provinciis Florentina et Pisana” (destroyed?).

  • Cimbex vespiformis Olivier, 1790, Encyc. Méth.: 772, sex not stated –“en France, en Italie … aux environs de Paris” (destroyed?).

  • Masaris apiformis Fabricius, 1793, Entomol. Syst. 2: 284, sex not stated –“in Italia” (Copenhagen).

  • Celonites abbreviatus var. Hungaricus Mocsáry, 1877, Termesz. Füzetek. 1: 90, female –“Hungaria centrali” (Budapest?).

  • Celonites abbreviatus var. engadinensis von Schulthess, 1923, in Michaelsen, Beitr. Kennt. Land-u. Süsswasser. Deutsch-Südwestafr. 2(2), Hym. VI Vesp.: 139, male, female –“Schweiz. Alpengebiet, Ober-Engadin, Sils Baselgia” (Zürich).

  • Distribution: Germany; Austria; Hungary; Switzerland; France; Spain; Portugal; Italy; Croatia; Albania; Bulgaria; Greece: Corfu; Cyprus; Turkey; Israel; Morocco.

  • abbreviatus tauricus Kostylev

  • Celonites abbreviatus tauricus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 108, female (in subgenus Celonites) – i0003-0082-3325-1-1-eq1.gif Crimea] (St. Petersburg?).

  • Distribution: Ukraine.

  • afer Lepeletier

  • Celonites afer Lepeletier, 1841, Hist. Nat. Insect. Hym. 2: 585, female –“Oran” (Paris?).

  • Distribution: Morocco; Algeria; Tunisia; Libya.

  • andrei Brauns

  • Celonites Andréi Brauns, 1905, Ann. Mus. Natl. Hung. 3: 228, female –“Terra Capensis: Willowmore” (Pretoria).

  • Distribution: South Africa: Western Cape, Eastern Cape.

  • bergenwahliae Gess

  • Celonites bergenwahliae Gess, 1989, Ann. Cape Prov. Mus. (Nat. Hist.) 18: 83, figs. 8–9, 12–13, 91, female, male –“Cape Province: Clanwilliam District, Klein Alexandershoek (32°20;pr20;dpS, 18°46;prE)” (holotype female, Grahamstown); also from 5 km W of Clanwilliam.

  • Distribution: South Africa: Western Cape.

  • capensis Brauns

  • Celonites capensis Brauns, 1905, Ann. Mus. Natl. Hung. 3: 231, female –“Willowmore” (Pretoria).

  • Distribution: South Africa: Western Cape, Eastern Cape.

  • clarus Gusenleitner

  • Celonites clarus Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 60, 66, female –“Iran, Teheran, Steppa presso Chitgar, 18 km W Teheran” (Venice).

  • Distribution: Iran.

  • clypeatus Brauns

  • Celonites clypeatus Brauns, 1913, Entomol. Mitt., Berlin 2: 206, female –“Willowmore” (Pretoria).

  • Distribution: Namibia*; South Africa: Northern Cape*, Eastern Cape.

  • crenulatus Morawitz

  • Celonites crenulatus Morawitz, 1888, Horae Soc. Entomol. Ross. 22: 267, female –“in territorio transcaspico (Kiltitschinar)” (St. Petersburg?).

  • Distribution: Iran*; Turkmenistan.

  • cyprius cyprius de Saussure

  • Celonites cyprius de Saussure, 1854, Mém. Soc. Phys. Hist. Nat. Genève 14: 4, male –“l'Ille de Chypre” (Geneva?).

  • Distribution: Cyprus.

  • cyprius smyrnensis Richards

  • Celonites cyprius smyrnensis Richards, 1962, Revision. Study Masarid Wasps: 218 (key), 232, female, male –“Turkey: near Smyrna, Burnova” (holotype female, London); also from Israel (Greece: Rhodes mentioned in key).

  • Distribution: Greece: Thessalia, Pelopponisos, Rhodes; Turkey; Armenia; Iran; Syria; Lebanon; Israel.

  • davidi Gess

  • Celonites davidi Gess, 1989, Ann. Cape Prov. Mus. (Nat. Hist.) 18: 83, fig. 14, 92, male –“Cape Province: Namaqualand, Anenous (29°15′30″S, 17°34′45″E)” (Grahamstown).

  • Distribution: South Africa: Northern Cape.

  • discretus Gusenleitner

  • Celonites discretus Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 60, figs. 3a, 4b, 62, female –“Iran, Fars, Monti steppiti 5 km N di Persepolis” (Venice).

  • Distribution: Iran.

  • fischeri Spinola

  • Celonites fischeri Spinola, 1838, Ann. Soc. Entomol. France 7: 505, female –“Egypte” (type depository unknown).

  • ?Celonites savignyi de Saussure, 1855, Ét. Fam. Vesp. 3: 91 –“Description de l'Egypte. Hymen. Pl. IX, fig. 19, ♀♂” (type depository unknown).

  • Distribution: Algeria; Tunisia; Libya; Egypt; Cyprus; Israel; Saudi Arabia.

  • foveolatus foveolatus Kostylev

  • Celonites foveolatus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 111, 112, male, female (in subgenus Paracelonites) – i0003-0082-3325-1-1-eq2.gif [Zakaspian desert] (Moscow, St. Petersburg?).

  • Distribution: Turkmenistan.

  • foveolatus nigrior Richards

  • Celonites foveolatus nigrior Richards, 1962, Revision. Study Masarid Wasps: 218 (key), 234, female –“Israel: Jericho” (Oxford).

  • Distribution: Turkey; Israel.

  • gariepensis Gess

  • Celonites gariepensis Gess, 1997, J. Hym. Res. 6: 36, 41, figs. 1–6, female, male –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown); also from two other localities in Richtersveld.

  • Distribution: South Africa: Northern Cape.

  • guichardi Richards

  • Celonites guichardi Richards, 1962, Revision. Study Masarid Wasps: 218 (key), 233, female –“Libya: Fezzan, El Manakh (50 m. N.W. of Edri)” (London).

  • Distribution: Libya.

  • hamanni Gusenleitner

  • Celonites hamanni Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 60, 65, 66, fig. 6, female, male –“Türkei Mut” (holotype female, Linz).

  • Celonites hammani[!]; Yildirim and Özbek, 1997, Atatürk. Ü. Zir. Fak. Der. 28 (2): 224.

  • Distribution: Turkey; Syria; Jordan.

  • hellenicus Gusenleitner

  • Celonites hellenicus Gusenleitner, 1997, Linz. Biol. Beitr. 29(1): 109, 110 (key), 112, male, female (in subgenus Eucelonites) –“Hellas: Ano Karmes, (Likeo Bgl.)” (holotype male, BOL); also from elsewhere in Greece.

  • Distribution: Greece.

  • humeralis Richards

  • Celonites humeralis Richards, 1962, Revision. Study Masarid Wasps: 219, 236, female –“Cape Province: near Calvinia, Bloukrans” (London); also from Nieuwoudtville.

  • Distribution: South Africa: Northern Cape.

  • hystrix Kostylev

  • Celonites hystrix Kostylev, 1940, Bull. Soc. Nat. Moscou (Biol.) 49: 138, female, male (in subgenus Eucelonites) –“Kondara, Chaîne Ghissar, 30 km vers le N de Stalinabad (autrefois Duchambé), 1 100 m au –d. du niv. de la mer” (coll. Gussakovskii, St. Petersburg).

  • Distribution: Tajikistan.

  • jousseaumei jousseaumei du Buysson

  • Celonites jousseaumei du Buysson, 1906, Rev. Entomol., Caen 25: 104, female, male –“Obock, Djibouti, Tadjourah” (Paris).

  • Celonites Joussaumei[!]; von Schulthess, 1923, in Michaelsen, Beitr. Kennt. Land-u. Süsswasser. Deutsch-Südwestafr. 2(2), Hym. VI Vesp.: 140.

  • Celonites jousseaumei asrensis Giordani Soika, 1957, Brit. Mus. Exped. S.-W. Arabia 1(31): 472, female –“YEMEN: ‘Asr, about 3 miles west of San'a, 8100 ft.” (London).

  • Distribution: Algeria; Sudan; Dijibouti; Israel; Saudi Arabia; Yemen; Oman; United Arab Emirates; Qatar.

  • jousseaumei senegalensis Richards

  • Celonites jousseaumei senegalensis Richards, 1962, Revision. Study Masarid Wasps: 216 (key), 224, female, male –“Senegal: Bambey” (holotype female, Paris).

  • Distribution: Senegal; Mali; Djibouti.

  • kostylevi Panfilov

  • Celonites kostylevi Panfilov, 1961, Arch. Mus. Zool. Univ. Moscou 8: 165, female, male (in subgenus Celonites) – i0003-0082-3325-1-1-eq3.gif [Kirghizia: east bank of Issyk-Kul’ lake, forest valley Ak-Ulen] (Moscow).

  • Distribution: Kyrgyzstan.

  • kozlovi Kostylev

  • Celonites kozlovi Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 110 (in subgenus Paracelonites) – i0003-0082-3325-1-1-eq4.gif [Gobi Desert, Mongolia] (St. Petersburg?).

  • Distribution: Mongolia.

  • laetus Panfilov

  • Celonites laetus Panfilov, 1968, Arch. Zool. Mus. Moscow State Univ. 11: 40, fig. 3, female – i0003-0082-3325-1-1-eq5.gif [Transcaucasia: Nakhichevan’, vicinity of Dzhulfa, desert foothills] (Moscow).

  • Distribution: Azerbaijan.

  • latitarsis Gess

  • Celonites latitarsis Gess, 1992, J. Hym. Res. 1: 141, figs. 1–5, female, male –“Cape Province: Clanwilliam District, 11 km W of Clanwilliam on road to Graafwater” (holotype female, Grahamstown).

  • Distribution: South Africa: Western Cape.

  • lobeliae Gess

  • Celonites lobeliae Gess, 1997, J. Hym. Res. 6: 36, 46, figs. 13–14, female, male –“Cape Province: Betw[een] Nieuwoudtville and top of Vanrhyns Pass” (holotype female, Grahamstown).

  • Distribution: South Africa: Northern Cape.

  • longipilis Gusenleitner

  • Celonites longipilis Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 59, figs. 3b, 4a, female –“Iran, Fers [Fars], Monti steppici 5 km N di Persepolis” (Venice).

  • Distribution: Iran.

  • mayeti Richards

  • Celonites mayeti Richards, 1962, Revision. Study Masarid Wasps: 222 (key), 243, figs. 202, 208, 211, female, male –“France: Bouches-du-Rhône, Camargue” (holotype male, Paris); also from other localities in France and Spain.

  • Distribution: France; Spain.

  • michaelseni von Schulthess

  • Celonites Michaelseni von Schulthess, 1923, in Michaelsen, Beitr. Kennt. Land.-u. Süsswasser. Deutsch-Südwestafr. 2(2), Hym. VI Vesp.: 137, male –“Windhuk” (Hamburg?).

  • Distribution: Namibia.

  • modestus biinterruptus Kostylev

  • Celonites modestus biinterruptus Kostylev, 1940, Bull. Soc. Nat. Moscou (Biol.) 49: 138, female, male (in subgenus Paracelonites) –“Ruidacht, 40 km. de Stalinabad” (coll. Gussakovskii, St. Petersburg).

  • Celonites modestus bisinterruptus[!]; Richards, 1962, Revision. Study Masarid Wasps: 247.

  • Distribution: Tajikistan.

  • modestus modestus Kostylev

  • Celonites modestus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 110 (in subgenus Paracelonites) – i0003-0082-3325-1-1-eq6.gif [Pamir, Mountains of Peter the Great] (St. Petersburg?).

  • Distribution: Tajikistan.

  • montanus Mocsáry

  • Celonites montanus Mocsáry, 1906, Ann. Hist.-Nat. Mus. Hung. 4: 198, male –“Turkestan: Alai montes” (Budapest).

  • Distribution: Kyrgyzstan; China: Xinjiang.

  • nursei Dover

  • Celonites nursei Dover, 1925 (1924), J. Proc. Asiat. Soc. Bengal (n. ser.) 20(6): 289, fig. 1, female –“Quetta” (London).

  • Distribution: Pakistan.

  • octoannulatus hissaricus Kostylev

  • Celonites octoannulatus hissaricus Kostylev, 1940, Bull. Soc. Nat. Moscou (Biol.) 49: 137, female, male (in subgenus Paracelonites) –“Ruidacht, 40 km vers le N de Stalinabad (autrefois Duchambé). 3000 m. au-dessus du niv. de la m.” (coll. Gussakovskii, St. Petersburg).

  • Distribution: Tajikistan.

  • octoannulatus octoannulatus (Kuznetzov)

  • Saryara octoannulata Kuznetzov, 1923, J. Turkestan Branch Russ. Geogr. Soc. 16: 127, female, male –“Montes Ac-Tasch, distr. Taschkent, Turkestan. Alt 7500” (St. Petersburg).

  • Distribution: Uzbekistan.

  • osseus Morawitz

  • Celonites osseus Morawitz, 1888, Horae Soc. Entomol. Ross. 22: 268, female –“in territorio transcaspico (Tschikischljar)” (St. Petersburg?).

  • Distribution: Armenia; Turkmenistan.

  • peliostomi Gess

  • Celonites peliostomi Gess, 1989, Ann. Cape Prov. Mus. (Nat. Hist.) 18: 83, figs. 1–5, female, male –“Cape Province: Namaqualand, Springbok, Hester Malan Nature Reserve” (holotype female, Grahamstown).

  • Distribution: South Africa: Northern Cape.

  • persicus Richards

  • Celonites persicus Richards, 1962, Revision. Study Masarid Wasps: 221 (key), 241, female –“South West Persia: K. Sefid” (London).

  • Distribution: Iran.

  • phlomis Gusenleitner

  • Celonites phleomis[!]; Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 60, fig. 3c. Incorrect original spelling.

  • Celonites phlomis Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 61, 63, fig. figs. 4c, 5, female, male –“Türkei, Urfa” (holotype female, Linz).

  • Distribution: Turkey.

  • pictus pictus Richards

  • Celonites pictus Richards, 1962, Revision. Study Masarid Wasps: 217 (key), 226, figs. 176–177, male –“Senegal: Bambey” (Paris).

  • Distribution: Senegal; Morocco; Saudi Arabia; Oman.

  • pictus rufiventris Gusenleitner

  • Celonites pictus rufiventris Gusenleitner, 1992, Linz. Biol. Beitr. 24(1): 59, 60, female –“Iran, Bandar Abbas (Strand)” (Linz).

  • Distribution: Iran.

  • promontorii Brauns

  • Celonites promontorii Brauns, 1905, Ann. Mus. Natl. Hung. 3: 232, female –“Willowmore” (Pretoria).

  • Distribution: South Africa: Northern Cape, Eastern Cape, Free State.

  • purcelli Brauns

  • Celonites Purcelli Brauns, 1905, Ann. Mus. Natl. Hung. 3: 226, female, male –“Terra Capensis: Willowmore” (Pretoria).

  • Celonites purcelli var. flavomaculatus Brauns, 1913, Entomol. Mitt., Berlin 2: 205, male –“Willowmore” (Pretoria).

  • Distribution: South Africa: Eastern Cape.

  • rothschildi du Buysson

  • Celonites rothschildi du Buysson, 1906, Rev. Entomol., Caen 25: 105, male –“Afrique orientale anglaise: Lesammise, Rendilé” (Paris).

  • Celonites Rotschildi[!]; von Schulthess, 1923, in Michaelsen, Beitr. Kennt. Land.-u. Süsswasser. Deutsch-Südwestafr. 2(2), Hym. VI Vesp.: 140.

  • Distribution: Kenya; Iran?

  • rudesculptus Kostylev

  • Celonites rudesculptus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 109 (in subgenus Paracelonites) – i0003-0082-3325-1-1-eq7.gif [Armenia] (St. Petersburg?).

  • Distribution: Armenia.

  • rugiceps Bischoff

  • Celonites rugiceps Bischoff, 1928, Abhand. Naturwiss. Verein. Bremen 27 (1): 86, female –“Kreta” (Berlin); also from mainland Greece; Turkey.

  • Distribution: Croatia; Yugoslavia; Bulgaria; Greece: Argos, Crete; Cyprus; Turkey.

  • semenovi Kostylev

  • Celonites semenovi Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 112 (in subgenus Eucelonites) – i0003-0082-3325-1-1-eq8.gif [Persia] (St. Petersburg?).

  • Distribution: Iran.

  • spinosus Gusenleitner

  • Celonites spinosus Gusenleitner, 1966, Polskie Pismo Entomol. 36: 359, fig. 2, male, female –“Kusadasi” (holotype female, Linz); also from other localities in Turkey.

  • Celonites abbreviatus invitus Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 58, fig. 2b, female, male –“Türkei, Gürün” (holotype female, Linz); also from elsewhere in Turkey, and Armenia.

  • Distribution: Turkey; Armenia.

  • tristiculus karatauicus Kostylev

  • Celonites tristiculus karatauicus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 111, male (in subgenus Paracelonites) –[Kara Tau] (St. Petersburg?).

  • Distribution: Kazakhstan.

  • tristiculus tristiculus Kostylev

  • Celonites tristiculus Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 111, female, male (in subgenus Paracelonites) – i0003-0082-3325-1-1-eq9.gif [Fergana. Syr Darya] (Moscow, St. Petersburg?).

  • Distribution: Uzbekistan.

  • tumidiscutellatus Gess

  • Celonites tumidiscutellatus Gess, 1997, J. Hym. Res. 6: 36, 44, figs. 7–12, female, male –“Cape Province: Namaqualand, Springbok, Hester Malan [now Goegap] Nature Reserve” (holotype female, Grahamstown); also from three other localities in Namaqualand, and Eastern Cape.

  • Distribution: South Africa: Northern Cape, Eastern Cape.

  • turneri Richards

  • Celonites turneri Richards, 1962, Revision. Study Masarid Wasps: 221 (key), 238, figs. 195–197, male –“Cape Province: Aliwal North” (London).

  • Distribution: South Africa: Western Cape,* Eastern Cape.

  • varipennis Richards

  • Celonites varipennis Richards, 1962, Revision. Study Masarid Wasps: 219 (key), 239, fig. 200, female, male –“Libya: Tibesti, Enneri Meche” (holotype male, London).

  • Distribution: Libya.

  • wahlenbergiae Gess

  • Celonites wahlenbergiae Gess, 1989, Ann. Cape Prov. Mus. (Nat. Hist.) 18: 83, figs. 6–7, 10–11, 87, female, male –“Cape Province: Clanwilliam District, Clanwilliam Dam” (holotype female, Grahamstown).

  • Distribution: South Africa: Western Cape.

  • wheeleri Brauns

  • Celonites Wheeleri Brauns, 1905, Ann. Mus. Natl. Hung. 3: 230, female, male –“Terra Capensis: Willowmore” (Pretoria).

  • Celonites wheeleri var. immaculatus Brauns, 1905, Ann. Mus. Natl. Hung. 3: 230, female –“Terra Capensis: Willowmore” (Pretoria).

  • Distribution: Namibia*; South Africa: Eastern Cape.

  • yemenensis ethiopicus Richards

  • Celonites yemenensis ethiopicus Richards, 1962, Revision. Study Masarid Wasps: 217 (key), 228, female –“Ethiopia: Diredawa” (London).

  • Distribution: Ethiopia.

  • yemenensis yemenensis Giordani Soika

  • Celonites yemenensis Giordani Soika, 1957, Brit. Mus. Exped. S.-W. Arabia 1(31): 472, fig. 1, male, female –“YEMEN: Ta'izz, on road to Mocha, 4100 ft.” (holotype male, London).

  • Celonites yemenensis yemensis[!]; Richards, 1969, J. Nat. Hist. 3: 82.

  • Distribution: Saudi Arabia; Yemen.

  • zavattarii Giordani Soika

  • Celonites zavattarii Giordani Soika, 1944, Atti Inst. Ven. Sci. Lett. Art. 103(2): 149, female –“Etiopia: Asile” (lectotype female, Venice); also from Caschei.

  • Distribution: Ethiopia.

  • Genus Ceramiopsis Zavattari

  • Ceramiopsis Zavattari, 1910, Ann. Mus. Civ. Stor. Nat. Genova 44: 531, genus. Type species: Ceramiopsis gestroi Zavattari, 1910, by monotypy.

  • gestroi Zavattari

  • Ceramiopsis gestroi Zavattari, 1910, Ann. Mus. Civ. Stor. Nat. Genova 44: 533, figs. A, B, female –“Urucú, Corumbà (Matto Grosso)” (Genova).

  • Distribution: Bolivia: Santa Cruz; Brazil: Mato Grosso do Sul; Paraguay: Guairá, Paraguarí, San Pedro; Argentina: Formosa, Santa Fé, Entre Rios.

  • paraguayensis Bertoni

  • Ceramiopsis paraguayensis Bertoni, 1921, Rev. Soc. Cient. Paraguay 1: 11, female, male –“Asunción, ‘Quinta Caballero’” (San Lorenzo).

  • Distribution: Bolivia: Santa Cruz; Paraguay: Alto Paraguay, Presidente Hayes, Central, Paraguarí; Argentina: Formosa.

  • Genus Ceramius Latreille

  • Ceramius Latreille, 1810, Consid. Gén. Crust. Arach. Insect.: 329, 438. Type species: Ceramius fonscolombei Latreille, 1810, by original designation.

  • Gnatho Klug, 1810, Mag. Ges. Naturf. Fr. Berl. 4: 32, 36, genus. Non Illiger, 1807 (Coleoptera). Type species: Gnatho lichtensteinii Klug, 1810, by monotypy.

  • Ceramias[!] Westwood, 1840, Introd. Modern Class. Insect. 2: 237.

  • Ceramus[!] Westwood, 1840, Introd. Modern Class. Insect. 2: 243.

  • Paraceramius de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III; 1855, Ét. Fam. Vesp. 3: 67, division of genus Ceramius Latreille (4 species). Type species: Ceramius spiricornis de Saussure, 1854, by subsequent designation of Ashmead, 1902, Can. Entomol. 34: 220.

  • Ceramioides de Saussure, 1854, Ét. Fam. Vesp. 3: pl. IV no. 1; 1855, Ét. Fam. Vesp. 3: 71, division of genus Ceramius Latreille; 1855: 71, division III of genus Ceramius. Type species: Ceramius cerceriformis de Saussure, 1854, by monotypy.

  • Euceramius Dalla Torre, 1904, Genera Insect. 19: 5, name for division I of Ceramius Latreille in de Saussure, 1854, Ét. Fam. Vesp. 3: pl. II, 1855, Ét. Fam. Vesp. 3: 62 (3 species). Type species: Ceramius fonscolombei Latreille, by subsequent designation of van der Vecht and Carpenter, 1990, Zool. Verh., Leiden 260.

  • auctus (Fabricius)

  • Liris aucta Fabricius, 1804, Syst. Piezat.: 229, sex not stated –“in Tangier” (female Copenhagen).

  • Ceramius spiricornis de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xx. Nomen nudum.

  • Ceramius spiricornis de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III, fig. 5, male; 1855, Ét. Fam. Vesp. 3: 68 –“L'Europe méridionale, la France, l'Espagne” (Paris).

  • Distribution: Morocco; Algeria.

  • beaumonti (Giordani Soika)

  • Paraceramius beaumonti Giordani Soika, 1957, Bull. Soc. Sci. Nat. Phys. Maroc 37: 168, figs. 1, 3, 5, male, female –“Asni” (holotype male, Venice); also from elsewhere in Morocco.

  • Distribution: Algeria; Morocco.

  • beyeri Brauns

  • Ceramius Beyeri Brauns, 1903, Z. Syst. Hym. Dipt. 3(2): 69, female, male –“Willowmore … Steytlerville in der Karroo des Caplandes” (Pretoria).

  • Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape.

  • bicolor (Thunberg)

  • Philanthus bicolor Thunberg, 1815, Nova Acta Reg. Soc. Sci. Uppsala 7: 127, 131, 289, sex not stated (male, Uppsala).

  • Ceramius Karrooensis Brauns, 1902, Z. Syst. Hym. Dipt. 2(5–6): 282, female, male –“Willowmore im Capland” (Pretoria).

  • Ceramius karooensis[!]; Brauns, 1910, Z. Wiss. Insektenbiol. 6(11): 387.

  • Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape.

  • bischoffi Richards

  • Ceramius bischoffi Richards, 1963, Zool. Meded. 38: 216, fig. 2, male, female –“Spain: Tornavacas, prov. Cáceres, 1200 m” (holotype male, Leiden); also from elsewhere in Spain.

  • Distribution: Spain.

  • braunsi Turner

  • Ceramius braunsi Turner, 1935, Ann. Mag. Nat. Hist. (10) 15: 291 (key), 294, fig. 2, male, female –“Olifant's River, between Citrusdal and Clanwilliam” (holotype male, Cape Town).

  • Distribution: South Africa: Western Cape, Eastern Cape.

  • brevitarsis Gess

  • Ceramius brevitarsis Gess, 1997, J. Hym. Res. 6: 36, 37, female, male –“Cape Province: Namaqualand, Richtersveld N[ational] Park, bet[ween] hills (28.10S, 17.02E), NW Koeroegabvlakte” (holotype female, Grahamstown).

  • Distribution: South Africa: Northern Cape.

  • bureschi Atanassov

  • Ceramius bureschi Atanassov, 1938, Mitt. K. Naturwiss. Inst. Sofia 11: 114, figs. 1, 2a, 3a, 4a, 5a, 6a, male –“Bulgarien, im Park des Palais Wrana bei Sofia” (Sofia).

  • Ceramius bureschi lycaonius Blüthgen, 1952, Mitt. Münch. Entomol. Ges. 42: 18, male, female –“Attika” (holotype male, Berlin).

  • Distribution: Bulgaria; Greece; Turkey.

  • caffer de Saussure

  • Ceramius caffer de Saussure, 1855, Ét. Fam. Vesp. 3: 76, female –“Cap de Bonne-Espérance” (Torino).

  • Ceramius consobrinus de Saussure, 1855, Ét. Fam. Vesp. 3: 77, female –“Cap de Bonne-Espérance” (Torino).

  • Distribution: South Africa: Western Cape.

  • capensis de Saussure

  • Ceramius capensis de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III, figs. 3–3a, female; 1855, Ét. Fam. Vesp. 3: 66 –“Cap de Bonne-Espérance” (Turin?).

  • Distribution: South Africa: Western Cape.

  • capicola Brauns

  • Ceramius capicola Brauns, 1902, Z. Syst. Hym. Dipt. 2(5): 278, female, male –“Willowmore” [“Hier”] (Pretoria).

  • Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape, Free State.

  • caucasicus André

  • Ceramius Caucasicus André, 1884, Spec. Hym. Europe Algérie II: 820, male –“Caucase” (Paris?).

  • Distribution: Iran; Armenia; Turkey.

  • cerceriformis de Saussure

  • Ceramius cerceriformis de Saussure, 1853, Ann. Soc. Entomol. France (3), Bull. II: xxi, male –“Cap de Bonne-Espérance” (Torino).

  • Ceramius vespiformis de Saussure, 1855, Ét. Fam. Vesp. 3: 79, female –“Cap de Bonne-Espérance” (Torino).

  • Ceramius Schulthessi Brauns, 1902, Z. Syst. Hym. Dipt. 2 (3): 182, female –“Willowmore, Capland” (Pretoria).

  • Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape.

  • clypeatus Richards

  • Ceramius clypeatus Richards, 1962, Revision. Study Masarid Wasps: 83, 88 (key), 99, male –“Clan William” (Pretoria).

  • Distribution: South Africa: Western Cape.

  • damarinus Turner

  • Ceramius damarinus Turner, 1935, Ann. Mag. Nat. Hist. (10) 15: 292 (key), 293, male, female –“Ongandjera, S.W. Africa … Kamanyab” (holotype male, Cape Town).

  • Distribution: Namibia.

  • fonscolombei fonscolombei Latreille

  • Ceramius fonscolombei Latreille, 1810, Consid. Gén. Crust. Arach. Insect.: 438, sex not stated [male] –“aux environs d'Aix” [p. 329] (Paris).

  • Distribution: France; Spain; Portugal.

  • fonscolombei oraniensis Lepeletier

  • Ceramius Oraniensis Lepeletier, 1841, Hist. Nat. Insect. Hym. 2: 591, female, male –“Oran” (Paris).

  • Ceramius Doursii André, 1884, Spec. Hym. Europe Algérie II: 818, female, male –“Algérie” (type depository unknown).

  • Distribution: Morocco; Algeria.

  • gessi Mauss

  • Ceramius maroccanus gessi Mauss, 1998, Beitr. Hym.-Tag. Stuttgart (2–4.10.1998): 18. Nomen nudum.

  • Ceramius gessi Mauss, 1999, Entomol. Scand. 30: 323, 331 (key), 340, figs. 1c, 2c, 3c, 3f, i, 4c, 4g, 4j, 5d–f, 7b, 8c, 9c, 10–11, female, male –“Morocco, 10 km W Tiznit, 29°43;prN 09°50;prW” (holotype male, BOL); also from elsewhere in Morocco.

  • Distribution: Morocco.

  • hispanicus Dusmet

  • Ceramius hispanicus Dusmet, 1909, Act. Mem. Primer Congr. Nat. Españ., Zaragoza, 1908: 179, 180, female, male –“Madrid! y Montarco! (en Madrid). –Jaramiel! (en Valladolid)” (Madrid).

  • Distribution: Spain.

  • jacoti Richards

  • Ceramius nigripennis de Saussure; Brauns, 1913, Entomol. Mitt., Berlin 2: 201, pl. 2, fig. 3, male. Misidentification.

  • Ceramius jacoti Richards, 1962, Revision. Study Masarid Wasps: 83, 89 (key), 101, male, female –“Cape Province: Ladismith” (holotype male, London); also from Montagu.

  • Distribution: South Africa: Western Cape.

  • lichtensteinii (Klug)

  • Gnatho Lichtensteinii Klug, 1810, Mag. Ges. Naturf. Fr. Berlin 4: 38, pl. 1 figs. 3, e, f, female –“Cap” (Berlin?).

  • Ceramius macrocephalus de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III figs. 2, 2a, 2c, female; 1855, Ét. Fam. Vesp. 3: 76 –“Cap de Bonne-Espérance” (Paris).

  • Ceramius rufomaculatus Cameron, 1906, Trans. South African Philos. Soc. 16(4): 325, female –“Pearston, Cape Colony” (London).

  • Distribution: Namibia?; South Africa: Northern Cape, Western Cape, Eastern Cape, Free State.

  • linearis Klug

  • Ceramius linearis Klug, 1824, Entomol. Monogr.: 227, male –“Prom. bonae spei, Distr. Vitenhagen” (Berlin).

  • Ceramius fumipennis Brauns, 1902, Z. Syst. Hym. Dipt. 2(5): 275, female, male –“Karroosteppe und … Port Elizabeth nahe der Küste” (Pretoria).

  • Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape.

  • lusitanicus Klug

  • Ceramius lusitanicus Klug, 1824, Entomol. Monogr.: 223, 230, sex not stated –“Lusitania” (Berlin).

  • Ceramius lusitanicus var. luteo-clypeata Dusmet, 1909, Act. Mem. Primer Congr. Nat. Españ., Zaragoza, 1908: 179, 180, male –“Montarco! (en Madrid)” (Madrid).

  • Distribution: Spain; Portugal; Gibraltar.

  • maroccanus (Giordani Soika)

  • Paraceramius maroccanus Giordani Soika, 1957, Bull. Soc. Sci. Nat. Phys. Maroc 37: 170, figs. 6, 8, 11–12, female, male –“Mogador” (holotype male, Venice); also from elsewhere in Morocco.

  • Distribution: Morocco.

  • metanotalis Richards

  • Ceramius metanotalis Richards, 1962, Revision. Study Masarid Wasps: 83, 86 (key), 106, fig. 114, female –“Cape Province: Calvinia” (London); also from Nieuwoudtville.

  • Distribution: South Africa: Northern Cape, Western Cape.

  • micheneri Gess

  • Ceramius micheneri Gess, 1968, Novos Taxa Entomol. 57: 3, figs. 1–6, male, female –“Cape Province: Citrusdal” (holotype male, Cape Town); also from Olifants River between Klawer and Clanwilliam.

  • Distribution: South Africa: Western Cape.

  • montanus Gusenleitner

  • Ceramius montanus Gusenleitner, 1990, Linz. Biol. Beitr. 22(2): 565, 567, fig. 2, male, female –“Marokko, Hoher Atlas, Oukaimeden, 2600–2800 m” (Linz); also from elsewhere in Morocco.

  • Distribution: Morocco.

  • nigripennis de Saussure

  • Ceramius nigripennis de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III figs. 4, 4c, female; 1855, Ét. Fam. Vesp. 3: 69 –“Cap de Bonne-Espérance” (Torino).

  • Ceramius hessei Turner, 1935, Ann. Mag. Nat. Hist. (10) 15: 291 (key), 296, fig. 1, male, female –“Kamieskroon, Namaqualand” (holotype male, Cape Town).

  • Distribution: South Africa: Northern Cape, Western Cape.

  • palaestinensis (Giordani Soika)

  • Paraceramius palaestinensis Giordani Soika, 1957, Bull. Soc. Sci. Nat. Phys. Maroc 37: 172, female –“Palestine: Ein Geb” (Venice).

  • Ceramius palestinensis[!]; Yildirim and Özbek, 1997, Atatürk. Ü. Zir. Fak. Der. 28 (2): 225; Yildirim and Kojima, 1999, Nat. Hist. Bull. Ibaraki Univ. 3: 20.

  • Distribution: Israel; Syria; Turkey.

  • peringueyi Brauns

  • Ceramius Péringueyi Brauns, 1913, Entomol. Mitt., Berlin 2: 194, female –“in der Nähe der Capstadt” (Pretoria).

  • Distribution: South Africa: Western Cape.

  • rex de Saussure

  • Ceramius rex de Saussure, 1855, Ét. Fam. Vesp. 3: 75, female –“Cap de Bonne-Espérance” (Turin?).

  • Distribution: South Africa: Western Cape, Northern Cape.

  • richardsi Gess

  • Ceramius peringueyi Brauns; Richards, 1962, Revision. Stud. Masarid. Wasps: 100, male. Misidentification.

  • Ceramius richardsi Gess, 1965, Ann. S. Afr. Mus. 48: 219, 221, male, female –“Cape Province: Clanwilliam” (holotype male, Pretoria); also from Paleisheuwel.

  • Distribution: South Africa: Western Cape.

  • rubripes Gusenleitner

  • Ceramius maroccanus rubripes Gusenleitner, 1990, Linz. Biol. Beitr. 22(2): 565, 566, female –“Marokko, Anezal, Col de Tizin Bachoom, 1500 m, 7°3W 30°7N” (coll. H. Tussac); also from elsewhere in Morocco.

  • Distribution: Morocco.

  • socius Turner

  • Ceramius socius Turner, 1935, Ann. Mag. Nat. Hist. (10) 15: 291 (key), 297, male, female –“Worcester, Cape Province” (holotype male, London); also from elsewhere in Western Cape.

  • Distribution: South Africa: Western Cape.

  • toriger von Schulthess

  • Ceramius toriger von Schulthess, 1935, Ann. Mag. Nat. Hist. (10) 16: 383, female –“South Africa, Cape Colony … St. Rhynspass” (London); also from elsewhere in Northern Cape.

  • Distribution: South Africa: Northern Cape.

  • tuberculifer de Saussure

  • Ceramius tuberculifer de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xx. Nomen nudum.

  • Ceramius tuberculifer de Saussure, 1854, Ét. Fam. Vesp. 3: pl. III fig. 6, male; 1855, Ét. Fam. Vesp. 3: 69, as a synonym of lusitanicus Klug –“L'Espagne, le midi de la France” (Paris).

  • Ceramius lusitanicus Klug; de Saussure, 1855, Ét. Fam. Vesp. 3: 69, male, female (in part). Misidentification.

  • Distribution: France; Spain.

  • vechti Richards

  • Ceramius vechti Richards, 1963, Zool. Meded. 38: 213, fig. 1, male, female –“Spain: Tornavacas, prov. Cáceres, 1200 m” (holotype male, Leiden); also from elsewhere in Spain.

  • Distribution: Spain.

  • Genus Jugurtia de Saussure

  • Jugurtia de Saussure, July 1854, Ét. Fam. Vesp. 3: 19, 37, 42–44, generic and specific characters of J. numidica and J. oraniensis; pl. I fig. 4, antenna of J. oraniensis; genus. Type species: Celonites oraniensis Lepeletier, 1841, by subsequent designation of Ashmead, 1902, Can. Entomol. 34: 220.

  • Jugurthia[!] André, 1886, Spec. Hym. Europe Algérie II, Cat. Méthod. Syn.: 47.

  • Jugurthia Dalla Torre, 1894, Cat. Hym. 9: 5; E. Saunders, 1905, Trans. Entomol. Soc. London: 399. Unnecessary emendation of Jugurtia de Saussure, 1854.

  • Jujurtha[!] Ashmead, 1902, Can. Entomol. 34: 220.

  • Masariella Brauns, 1905, Ann. Mus. Natl. Hung. 3: 223, genus. Type species: Masariella alfkeni du Buysson, 1904, by monotypy.

  • Ceramiellus von Schulthess, 1922, Deutsch. Entomol. Z. 1922: 401, genus. Type species: Ceramiellus braunsi von Schulthess, 1922, by monotypy.

  • Juguartia[!] Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 68.

  • alfkeni (du Buysson)

  • Masaris alfkeni du Buysson, 1904, Bull. Soc. Entomol. France: 144, female –“Damara land, Swakopmund” (Bremen).

  • Distribution: Namibia.

  • algerica (von Schulthess)

  • Masariella? algerica von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 499 (key), 501, fig. 2, female –“Algeria occidentalis, Ain Sefra, … Ain Fezza” (Zürich).

  • Distribution: Morocco; Algeria; Libya.

  • biskrensis Bequaert

  • Jugurtia biskrensis Bequaert, 1937, Bull. Ann. Soc. Entomol. Belg. 68: 343, 345 (key), 358, fig. 5A–F, female, male –“Algérie. –Biskra” (holotype female, Cambridge).

  • Distribution: Morocco; Algeria.

  • braunsi (von Schulthess)

  • Ceramiellus braunsi von Schulthess, 1922, Deutsch. Entomol. Z. 1922: 402, 2 figs., female, male –“Südwestafrika: Groß- und Klein-Namaland, Grootfontein … Steinkopf” (Berlin).

  • Distribution: Namibia; South Africa: Western Cape, Eastern Cape.

  • braunsiella (von Schulthess)

  • Masaris braunsiella von Schulthess, 1930, Ann. Mag. Nat. Hist. (10) 5: 326, fig. 1, male –“Cape Province, Matjesfontein” (London).

  • Distribution: Namibia; South Africa: Northern Cape, Western Cape, Eastern Cape.

  • calcarata Richards

  • Jugurtia calcarata Richards, 1962, Revision. Study Masarid Wasps: 251 (key), 260, male, female –“S. Africa, Cape Province: Worcester” (holotype male, London); also from elsewhere in Western Cape.

  • Distribution: South Africa: Western Cape.

  • codoni Gess

  • Jugurtia codoni Gess, 1997, J. Hym. Res. 6: 36, 53, female, male –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown); also from three other localities in Richtersveld.

  • Distribution: South Africa: Northern Cape.

  • confusa Richards

  • Jugurtia confusa Richards, 1962, Revision. Study Masarid Wasps: 249, 251 (key), 258, figs. 213–215, 218, 220–221, female, male –“S. Africa: Cape Province, Willowmore” (holotype male, London).

  • Distribution: South Africa: Eastern Cape.

  • discrepans (Brauns)

  • Masaris discrepans Brauns, 1913, Entomol. Mitt., Berlin 2: 203, pl. II fig. 9, female, male –“Worcester im Süden der Kapkolonie … und bei Oudtshoorn (Kapland)” (Pretoria).

  • Distribution: Namibia; South Africa: Western Cape, Eastern Cape.

  • dispar (Dufour)

  • Celonites dispar Dufour, 1851, Ann. Soc. Entomol. France (2) 9: 58, 59, pl. 3 figs. 1–8, female, male –“environs de Madrid” (Paris?).

  • Distribution: Spain; Portugal; Gibraltar.

  • duplicata Richards

  • Jugurtia duplicata Richards, 1962, Revision. Study Masarid Wasps: 249, 254 (key), 263, male –“Cape Province: Van Rhynsdorp” (Pretoria).

  • Distribution: South Africa: Western Cape.

  • eburnea (Turner)

  • Masariella turneri eburnea Turner, 1935, Ann. Mag. Nat. Hist. (10) 15: 299, fig. 3, female, male –“Kamieskroon, Namaqualand” (holotype male, Cape Town).

  • Jugurtia polita Richards, 1962, Revision. Study Masarid Wasps: 251 (key), 256, female, male –“S. Africa, Cape Province: Calvinia” (holotype male, London); also from elsewhere in Cape Province.

  • Distribution: South Africa: Northern Cape.

  • escalerae Meade-Waldo

  • Jugurtia escalerae Meade-Waldo, 1910, Ann. Mag. Nat. Hist. (8) 5: 33, female –“S.W. Persia” (London).

  • Distribution: Turkey; Iran.

  • eurycara Kostylev

  • Jugurtia eurycara Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 94 (key), 97, figs. 6–7, male, female –“Caucasus, Armen.” (St. Petersburg); also from Erivanj [Yerevan].

  • Distribution: Turkey; Iran; Armenia.

  • irana Kostylev

  • Jugurtia irana Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 94 (key), 99, female, male –“Via Karvandar-Bampur, S.-O. Persien … Tshanshen-Sadk, Bampur, S.-O. Perasien … Kuusha-Larumba, Bampur, S.-O. Persien … Kurljut-Langer, Chorasan, O. Persien” (St. Petersburg).

  • Distribution: Iran.

  • jemenensis Kostylev

  • Jugurtia jemenensis Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 94 (key), 99, female –“Jemen, Sanaa” (Moscow).

  • Jugurtia hoggarica Giordani Soika, 1954, Boll. Mus. Civ. Stor. Nat. Venezia 7: 20, male –“Hoggar, Atakor, Guelta Afilale” (Venice).

  • Jugurtia jemenesis[!]; Richards, 1962, Revision. Study Masarid Wasps: 265.

  • Distribution: Algeria; Israel; Saudi Arabia; Yemen; Oman; United Arab Emirates.

  • koeroegabensis Gess

  • Jugurtia koeroegabensis Gess, 1997, J. Hym. Res. 6: 36, 55, female, male –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown); also from one other locality in Richtersveld.

  • Distribution: South Africa: Northern Cape.

  • nadigorum Bequaert

  • Jugurtia nadigorum Bequaert, 1937, Bull. Ann. Soc. Entomol. Belg. 68: 343, 344 (key), 347, fig. 2A–F, female, male –“Maroc. –Marrakech” (holotype female, Cambridge); also from one other locality.

  • Distribution: Morocco.

  • oraniensis (Lepeletier)

  • Celonites Oraniensis Lepeletier, 1841, Hist. Nat. Insect. Hym. 2: 586, female, male –“Oran” (Paris?).

  • Jugurtia numidica de Saussure, 1854, Ét. Fam. Vesp. 3: pl. V fig. 3, male; 1855, Ét. Fam. Vesp. 3: 85 –“L'Algérie” (Paris).

  • Jugurtia numida[!]; de Saussure, 1855, Ét. Fam. Vesp. 3: 85.

  • Distribution: Morocco; Algeria; Tunisia.

  • saussurei (Brauns)

  • Masaris Saussurei Brauns, 1905, Ann. Mus. Natl. Hung. 3: 219, female, male –“Terra Capensis: Willowmore” (Pretoria).

  • Distribution: South Africa: Western Cape, Eastern Cape.

  • simpsoni Meade-Waldo

  • Jugurtia simpsoni Meade Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 448, female –“Patchari … N'jau” (London).

  • Masariella? testaceopicta von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 499 (key), 502, fig. 3, female –“N. Nigeria, Zungeru” (London).

  • Distribution: Gambia; Senegal; Burkina Faso; Nigeria.

  • spinolae (de Saussure)

  • Masaris spinolae de Saussure, 1855, Ét. Fam. Vesp. 3: 95, female –“Cap de Bonne-Espérance” (Torino).

  • Jugurtia spinolai[!]; Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 92.

  • Jugurtia spirolai[!]; Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 100.

  • Distribution: South Africa: Western Cape.

  • tigrina Gess

  • Jugurtia tigrina Gess, 1997, J. Hym. Res. 6: 36, 50, figs. 1–6, female, male –“Cape Province: Namaqualand, Springbok, Hester Malan [now Goegap] Nature Reserve” (holotype female, Grahamstown); also from five other localities in Northern Cape.

  • Distribution: South Africa: Northern Cape.

  • turneri (von Schulthess)

  • Masariella? turneri von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 499 (key), 500, fig. 1, female –“Cape Province, Little Carroo, 38 m. E. of Ceres” (London).

  • Distribution: South Africa: Western Cape.

  • zarudnyi Kostylev

  • Jugurtia zarudnyi Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 93 (key), 94, figs. 4–5, male, female –“Chorasan … Kermän, Prov. Mokran” (St. Petersburg).

  • Distribution: Iran.

  • Genus Masarina Richards

  • Masarina Richards, 1962, Revision. Study Masarid Wasps: vii, 213, 268, genus. Type species: Masarina familiaris Richards, 1962, by original designation.

  • ceres Gess

  • Masarina ceres Gess, 1997, J. Hym. Res. 6: 36, 60, male, figs. 23–24 –“Cape Province: 17 km N of Ceres, near top of Gydo Pass” (Grahamstown).

  • Distribution: South Africa: Western Cape.

  • familiaris Richards

  • Masariella spinolae (de Saussure); Bequaert, 1928, Ann. Mag. Nat. Hist. (10) 2: 187. Misidentification.

  • Masarina familiaris Richards, 1962, Revision. Study Masarid Wasps: 268, 270, figs. 225–231, male, female –“Cape Province: Camps Bay” (holotype male, London); also from elsewhere in Cape Province.

  • Masarina communis; Richards, 1962, Revision. Study Masarid Wasps: 269. Lapsus calami.

  • Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape.

  • hyalinipennis Richards

  • Masarina hyalinipennis Richards, 1962, Revision. Study Masarid Wasps: 268, male –“Cape Province: van Rhynsdorp” (Pretoria).

  • Distribution: South Africa: Northern Cape, Western Cape.

  • mixta Richards

  • Masarina mixta Richards, 1962, Revision. Study Masarid Wasps: 268 (key), 271, female –“S. Africa, Cape Province: Nieuwoudtville” (London).

  • Distribution: South Africa: Northern Cape, Western Cape.

  • mixtoides Gess

  • Masarina mixtoides Gess, 1997, J. Hym. Res. 6: 36, 61, female, male, fig. 32 –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown); also from two other localities in Richtersveld.

  • Distribution: South Africa: Northern Cape.

  • namaqua Gess

  • Masarina namaqua Gess, 1997, J. Hym. Res. 6: 36, 65, female, male, figs. 25–27, 33 –“Cape Province: Namaqualand, Springbok, Hester Malan [now Goegap] Nature Reserve” (holotype female, Grahamstown); also from three other localities in Namaqualand.

  • Distribution: South Africa: Northern Cape.

  • parvula Gess

  • Masarina parvula Gess, 1997, J. Hym. Res. 6: 36, 65, female, male, figs. 28–30, 34 –“Cape Province: Namaqualand, Springbok, Hester Malan [now Goegap] Nature Reserve” (holotype female, Grahamstown); also from two other localities in Namaqualand.

  • Distribution: South Africa: Northern Cape.

  • peliostomi Gess

  • Masarina peliostomi Gess, 1997, J. Hym. Res. 6: 36, 67, female, male, figs. 35–39 –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown).

  • Distribution: South Africa: Northern Cape.

  • strucki Gess

  • Masarina strucki Gess, 1988, Ann. Cape Prov. Mus. (Nat. Hist.) 16: 351, 352, female –“[Cape Province:] Namaqualand, [Springbok]” (Grahamstown).

  • Distribution: South Africa: Northern Cape; Western Cape.

  • tylecodoni Gess

  • Masarina tylecodoni Gess, 1997, J. Hym. Res. 6: 36, 69, female, male, figs. 40–43 –“Cape Province: Namaqualand, Richtersveld National Park, Koeroegabvlakte (28.11S, 17.03E)” (holotype female, Grahamstown).

  • Distribution: South Africa: Northern Cape.

  • Genus Masaris Fabricius

  • Masaris Fabricius, 1793, Entomol. Syst. II: vi, 283, 284, genus (2 species). Type species: Masaris vespiformis Fabricius, 1793, by subsequent monotypy; Latreille, 1802, Hist. Nat. Crust. Insect. 3: 368, placed the second species in a new genus, Celonites.

  • Massaris Billberg, 1820, Enum. Insect.: 109. Unnecessary emendation of Masaris Fabricius.

  • Mazaris Schulz, 1911, Zool. Ann. Würzburg 4: 199. Unnecessary emendation of Masaris Fabricius.

  • Saryara Kuznetzov, 1923, J. Turkestan Branch Russ. Geograph. Soc. 16: 127, genus (2 species). Type species: Saryara longicornis Kuznetzov, 1923, by subsequent designation of van der Vecht and Carpenter, 1990, Zool. Verh., Leiden 260.

  • aegyptiacus aegyptiacus Meade-Waldo

  • Masaris vespiformis aegyptiacus Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 8: 447, fig. 2, male –“Cairo, Egypt” (London).

  • Distribution: Egypt; Israel; Jordan.

  • aegyptiacus arabicus Giordani Soika

  • Masaris vespiformis arabicus Giordani Soika, 1957, Brit. Mus. Exped. S.-W. Arabia 1(31): 472, female –“SAUDI ARABIA: Jebel Shammar Province, Hâil district” (London).

  • Distribution: Saudi Arabia.

  • carli von Schulthess

  • Masaris saussurei Carl, 1921, Rev. Suisse Zool. 28: 41, male, female –“Zarawchan; … Sutkend. Turkestan” (Geneva). Non Brauns, 1905.

  • Masaris carli von Schulthess, 1922, Deutsch. Entomol. Z. 1922: 404, replacement name for saussurei Carl.

  • Masaris smirnovi Kostylev, 1925, Entomol. Mitt. 14: 150 (key), 152, fig. 2, male –“Perovsk (Turkestan, Syr-Darjja-Gebiet)” (Moscow).

  • Distribution: Iran; Kazahkstan; Uzbekistan.

  • gussakovskii Kostylev

  • Masaris gussakovskii Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 87, 89, fig. 2, female –“Turkestan, distr. Chodzhent” (coll. Gussakovskii, St. Petersburg); also from Konstantinovsk.

  • Distribution: Uzbekistan.

  • longicornis (Kuznetzov)

  • Saryara longicornis Kuznetzov, 1923, J. Turkestan Branch Russ. Geogr. Soc. 16: 127, male –“Montes Ac-Tasch. Alt. 8000′” (St. Petersburg).

  • Distribution: Afghanistan; Uzbekistan; Kyrgyzstan.

  • tianshanicus Panfilov

  • Masaris tianshanicus Panfilov, 1968, Arch. Zool. Mus. Moscow State Univ. 11: 38, fig. 2, female – i0003-0082-3325-1-1-eq10.gif [Tian Shan: south bank of Issyk-Kul’ lake, Smaller Dzhargylchak (near Akterek), dry hills with steppe vegetation] (Moscow).

  • Distribution: Kyrgyzstan.

  • vespiformis Fabricius

  • Masaris vespiformis Fabricius, 1793, Entomol. Syst. 2: 283, sex not stated –“in Barbariae floribus” (Copenhagen).

  • Erynnis romandi de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xix, male –“Barbaria et Ægypta” (Paris).

  • Ceramius hylaeiformis Dalla Torre, 1894, Cat. Hym. 9: 4, as a synonym of vespiformis, attributed to Klug.

  • Distribution: Morocco; Algeria; Egypt.

  • Genus Pseudomasaris Ashmead

  • Pseudomasaris Ashmead, 1902, Can. Entomol. 34: 221, genus. Type species: Masaris occidentalis Cresson, 1871, by original designation.

  • Toryna Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 407, 411, subgenus of Pseudomasaris Ashmead. Type species: Masaris vespoides Cresson, 1863, by original designation and monotypy.

  • Holopticus Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 408, 414, subgenus of Pseudomasaris Ashmead. Type species: Masaris texanus Cresson, 1871, by original designation.

  • Cotyledon Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 408, 430, subgenus of Pseudomasaris Ashmead. Type species: Masaris edwardsii Cresson, 1872, by original designation.

  • basirufus Rohwer

  • Pseudomasaris zonalis basirufus Rohwer, 1912, Proc. U.S. Natl. Mus. 41: 452, female –“Death Valley, California” (Washington).

  • Pseudomasaris bariscipus Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 409 (key), 418, figs. 77, 85, male –“ARIZONA: Quartzite” (Philadelphia).

  • Pseudomasaris bariscapus[!]; Bequaert, 1940, Bull. Brooklyn Entomol. Soc. 35: 38.

  • Distribution: USA: California, Nevada, Arizona.

  • cazieri Bohart

  • Pseudomasaris cazieri Bohart, 1963, in Richards, Univ. Calif. Publ. Entomol. 27: 289 (key), 290, male –“Palos Colorados, Durango, Mexico, 8,000 feet elevation” (New York).

  • Distribution: Mexico: Durango, Chihuahua.

  • coquilletti Rohwer

  • Pseudomasaris coquilletti Rohwer, 1911, Proc. U.S. Natl. Mus. 40: 555, female, male –“Los Angeles County, California” (holotype female, Washington).

  • Distribution: USA: Oregon, California, Utah, Arizona.

  • edwardsii (Cresson)

  • Masaris edwardsii Cresson, 1872, Trans. Am. Entomol. Soc. 4: 87, female, male –“California” (Philadelphia).

  • Distribution: USA: Washington, Oregon, Idaho, Wyoming, Colorado, Utah, Nevada, California, Arizona; Mexico: Baja California Norte.

  • macneilli Bohart

  • Pseudomasaris macneilli Bohart, 1963, in Richards, Univ. Calif. Publ. Entomol. 27: 285 (key), 291, male, female –“Near Sonora Peak (above Sonora Pass), Tuolumne County, California, 11,000 feet elevation” (holotype male, San Francisco); also from elsewhere in California, and Utah.

  • Distribution: USA: California, Utah.

  • macswaini Bohart

  • Pseudomasaris macswaini Bohart, 1963, in Richards, Univ. Calif. Publ. Entomol. 27: 286 (key), 301, male, female –“Andreas Canyon, Riverside County, California” (holotype male, Davis); also from other localities in California.

  • Distribution: USA: California; Mexico*: Baja California Norte.

  • maculifrons (Fox)

  • Masaris maculifrons Fox, 1894, Proc. Calif. Acad. Sci. (2) 4: 12, female –“El Paraiso, L. Cal.” (San Francisco).

  • Pseudomasaris albifrons Rohwer, 1912, Proc. U.S. Natl. Mus. 41: 451, male –“New Mexico … Las Cruces” (Washington).

  • Pseudomasaris zonalis neomexicanus Rohwer, 1912, Proc. U.S. Natl. Mus. 41: 452, female –“Aztec, New Mexico” (Washington).

  • Pseudomasaris rohweri Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 409 (key), 417, male (in subgenus Holopticus) –“ARIZONA: Quartzite” (Philadelphia); also from Phoenix.

  • Distribution: USA: California, Nevada, Utah, Arizona, New Mexico; Mexico: Sonora, Baja California Norte.

  • marginalis (Cresson)

  • Masaris marginalis Cresson, 1864, Proc. Entomol. Soc. Philadelphia 3: 677, female –“Rocky Mountains, Colorado Territory” (Philadelphia).

  • Distribution: Canada: British Columbia, Alberta.; USA: Washington, Idaho, Wyoming, Colorado, Nevada, Utah, California, Arizona, New Mexico.

  • micheneri Bohart

  • Pseudomasaris micheneri Bohart, 1963, in Richards, Univ. Calif. Publ. Entomol. 27: 284, figs. 1, 14, 21, 286–287 (key), 298, male, female –“Westgard Pass, Inyo County, California, 6,000 feet elevation” (holotype male, Davis); also from elsewhere in California.

  • Distribution: USA: California, Nevada.

  • occidentalis (Cresson)

  • Masaris occidentalis Cresson, 1871, Trans. Am. Entomol. Soc. 3: 348, female –“Texas” (Philadelphia).

  • Distribution: USA: Kansas, Texas, New Mexico.

  • phaceliae Rohwer

  • Pseudomasaris phaceliae Rohwer, 1912, Proc. U.S. Natl. Mus. 41: 450, male, female –“New Mexico, Albuquerque” (holotype male, Washington); also from Mesilla and Filmore Canyon.

  • Distribution: USA: Texas, New Mexico, Arizona.

  • texanus (Cresson)

  • Masaris texanus Cresson, 1871, Trans. Am. Entomol. Soc. 3: 348, male, female –“Texas” (Philadelphia).

  • Distribution: USA: Texas.

  • vespoides (Cresson)

  • Masaris vespoides Cresson, 1863, Proc. Entomol. Soc. Philadelphia 2: 69, pl. 4, male, female –“Pike's Peak” (Philadelphia).

  • Pseudomasaris vespoides robertsoni Cockerell, 1913, Proc. Entomol. Soc. Wash. 15: 107, female –“Redlands, California” (Washington).

  • Distribution: USA: Nebraska, South Dakota, Montana, Washington, Oregon, Idaho, Wyoming, Colorado, Utah, Nevada, California, Arizona, New Mexico; Mexico: Baja California Norte.

  • wheeleri Bequaert

  • Pseudomasaris wheeleri Bequaert, 1929, Psyche 36: 61, fig. 1, male, female –“CALIFORNIA … Palm Springs, Riverside Co.” (holotype male, Cambridge).

  • Distribution: USA: California; Mexico: Baja California Norte.

  • zonalis (Cresson)

  • Masaris zonalis Cresson, 1864, Proc. Entomol. Soc. Philadelphia 3: 674, female, male –“Rocky Mountains, Colorado Territory” (Philadelphia).

  • Pseudomasaris zonalis albopictus Bohart, 1950, Proc. Biol. Soc. Wash. 63: 78, male, female –“Jenny Lake, Grand Teton National Park, Wyoming” (holotype male, San Francisco); also from five other localities in Idaho, Oregon and Washington.

  • Distribution: Canada: British Columbia; USA: Nebraska, Montana, Washington, Oregon, Idaho, Wyoming, Colorado, Utah, Nevada, California.

  • Genus Quartinia André

  • QuartiniaGribodo, in litteris” André, 1884, Spec. Hym. Europe Algérie II: 811, 822, genus. Type species: Quartinia dilecta André, by monotypy.

  • Juartinia[!] Cameron, 1904, Z. Syst. Hym. Dipt. 4: 89.

  • Quartiniella von Schulthess, 1929, Ann. Mag. Nat. Hist. 10(3): 504, 510, genus. Type species: Quartiniella waterstoni von Schulthess, 1929, by monotypy.

  • Quartinioides Richards, 1962, Revision. Study Masarid Wasps: 127, 170, genus. Type species: Quartinia capensis Kohl, 1898, by original designation.

  • affinis Richards

  • Quartinia affinis Richards, 1962, Revision. Study Masarid Wasps: 131 (key), 143, female –“Algeria: Sidi Farouch” (London).

  • Distribution: Algeria.

  • albopicta (Richards), new combination

  • Quartinoides[!] albopicta Richards, 1982, Boll. Mus. Civ. Stor. Nat. Venezia 32: 199, female –“S.W. Africa: Gobabeb, 25°36;prS, 15°10;prE” (Copenhagen).

  • Distribution: Namibia.

  • alcestis Richards

  • Quartinia alcestis Richards, 1962, Revision. Study Masarid Wasps: 133 (key), 164, female, male –“Cape Province: Worcester” (holotype female, London).

  • Distribution: South Africa: Western Cape.

  • andromeda (Richards), new combination

  • Quartinioides andromeda Richards, 1962, Revision. Study Masarid Wasps: 172 (key), 182, female –“Cape Province: Prince Albert Road” (London); also from Matjesfontein.

  • Distribution: South Africa: Western Cape.

  • antennata von Schulthess

  • Quartinia antennata von Schulthess, 1935, Ann. Mag. Nat. Hist. (10) 16: 385 (key), 388, fig., male –“South Africa, Cape Colony, Worcester” (London).

  • Distribution: South Africa: Western Cape.

  • antigone (Richards), new combination

  • Quartinioides antigone Richards, 1962, Revision. Study Masarid Wasps: 174 (key), 190, figs. 157–161, female, male –“Cape Province: Matjesfontein” (holotype male, London); also from elsewhere in Western Cape.

  • Distribution: South Africa: Western Cape.

  • araxana Giordani Soika

  • Quartinia araxana Giordani Soika, 1960, Boll. Soc. Entomol. Ital. 90: 133, female, male –“Caucaso: Araxesthal” (lectotype female, Venice).

  • Distribution: Armenia; Turkey; Iran.

  • arsinoe (Richards), new combination

  • Quartinioides arsinoe Richards, 1962, Revision. Study Masarid Wasps: 172 (key), 182, female –“S.W. Africa: Aus” (London); also from South Africa.

  • Distribution: Namibia; South Africa: Western Cape.

  • artemis Richards

  • Quartinia artemis Richards, 1962, Revision. Study Masarid Wasps: 128 (key), 156, female –“Cape Province: Calvinia” (London).

  • Distribution: South Africa: Northern Cape.

  • atra von Schulthess

  • Quartinia atra von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 505, female –“Cape Province, Montagu” (London).

  • Distribution: South Africa: Western Cape.

  • atrata Gusenleitner

  • Quartinia atrata Gusenleitner, 1998, Linz. Biol. Beitr. 30(2): 503, 511, female –“Iran, 15 km SE Sarvestan/Fars, 1800 m” (Linz).

  • Distribution: Iran.

  • basuto (Richards), new combination

  • Quartinioides basuto Richards, 1962, Revision. Study Masarid Wasps: 173 (key), 184, female, male –“Basutoland: Mamathes” (holotype male, London).

  • Distribution: Lesotho.

  • breyeri Richards

  • Quartinia breyeri Richards, 1962, Revision. Study Masarid Wasps: 130 (key), 157, female, male –“Cape Province: N.E. Zoutp. district” [actually Northern Transvaal] (holotype female, Pretoria).

  • Distribution: South Africa: Northern Transvaal.

  • canariensis Blüthgen

  • Quartinia canariensis Blüthgen, 1958, Comment. Biol. 18(6): 6, 11, female –“Fuerteventura: Matural” (Helsinki).

  • Distribution: Spain: Canary Islands: Fuerteventura.

  • candida Gusenleitner

  • Quartinia candida Gusenleitner, 1997, Linz. Biol. Beitr. 29(1): 137, 147, female –“Israel: 45 km N Elat, Sand tunes [dunes] E Qetura, 29°58′/35°06′” (coll. Schmid-Egger).

  • Distribution: Israel.

  • capensis Kohl

  • Quartinia capensis Kohl, 1898, Termesz. Füzetek 21: 365, female, pl. XV fig. 14 –“Capland … Algoa-Bai” (Vienna).

  • Quartinia scutellimacula von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 508, male, female –“Cape Province, Mossel Bay” (lectotype female, London); also from elsewhere in Western Cape.

  • Distribution: South Africa: Western Cape, Eastern Cape.

  • ceres (Richards), new combination

  • Quartinioides ceres Richards, 1962, Revision. Study Masarid Wasps: 177 (key), 206, female, male –“Cape Province: Little Karroo, 38 m. E. of Ceres” (holotype male, London); also from elsewhere in Cape Province.

  • Distribution: South Africa: Western Cape, Eastern Cape.

  • chlorotica (Morawitz)

  • Jugurtia chlorotica Morawitz, 1888, Horae Soc. Entomol. Ross. 22: 270, female –“in territorio transcaspico (Tschikischljar)” (St. Petersburg?).

  • Distribution: Turkmenistan.

  • cincta Benoist

  • Quartinia cincta Benoist, 1929, Bull. Soc. Sci. Nat. Maroc 9: 93, fig. 2, female, male –“Maroc: Asni, dans la vallée de la Reraya, … (altitude, 1.200 m.)” (Paris).

  • Distribution: Morocco.

  • cressida (Richards), new combination

  • Quartinioides cressida Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 203, female, male –“Cape Province: Camps Bay, Cape Peninsula” (holotype male, London); also from elsewhere in Cape Province, and Namibia.

  • Distribution: Namibia; South Africa: Northern Cape, Western Cape, Eastern Cape.

  • cretica Gusenleitner

  • Quartinia cretica Gusenleitner, 1994, Linz. Biol. Beitr. 26(1): 331, female –“Griechenland: Kreta, Kastro W Ammoudari; 35.18N 24.10E; 1000–1200 m” (Linz); also from elsewhere in Crete.

  • Distribution: Greece: Crete.

  • cyllene (Richards), new combination

  • Quartinioides cyllene Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 201, female, male –“Cape Province: 13 miles N.E. of Touws R.” (holotype female, London); also from Worcester.

  • Distribution: South Africa: Western Cape.

  • cynara (Richards), new combination

  • Quartinioides cynara Richards, 1962, Revision. Study Masarid Wasps: 173, 174 (key), 189, female –“Cape Province: Willowmore” (Pretoria); also from Prince Albert Road.

  • Distribution: South Africa: Western Cape, Eastern Cape.

  • diana (Richards), new combination

  • Quartinioides diana Richards, 1962, Revision. Study Masarid Wasps: 171 (key), 178, female –“S.W. Africa: Aus” (London).

  • Distribution: Namibia.

  • dilecta André

  • Quartinia DilectaGribodo (in litteris)” André, 1884, Spec. Hym. Europe Algérie II: 824, pl. XLV figs. 3, 7, female, male –“Tunis” (type depository unknown).

  • Distribution: Morocco; Algeria; Tunisia.

  • dryope (Richards), new combination

  • Quartinioides dryope Richards, 1962, Revision. Study Masarid Wasps: 173, 175 (key), 196, male –“Cape Province: Willowmore” (London).

  • Distribution: South Africa: Eastern Cape.

  • elissa (Richards), new combination

  • Quartinioides elissa Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 204, female, male –“Cape Province … Worcester” (holotype male, London); also from elsewhere in Western Cape.

  • Distribution: South Africa: Western Cape.

  • eremobia Richards

  • Quartinia eremobia Richards, 1962, Revision. Study Masarid Wasps: 135 (key), 137, female, male –“Algeria: 80 Km. nord d'El Goléa” (holotype male, Paris); also from Libya: Tripolitania.

  • Distribution: Algeria; Libya.

  • eurydice (Richards), new combination

  • Quartinioides eurydice Richards, 1962, Revision. Study Masarid Wasps: 172 (key), 183, female –“Cape Province: Bloukrans, near Calvinia” (London).

  • Distribution: South Africa: Northern Cape.

  • flava (Richards), new combination

  • Quartiniella flava Richards, 1962, Revision. Study Masarid Wasps: 208 (key), 210, female, male –“Cape Province: Nieuwoudtville” (holotype female, London).

  • Distribution: South Africa: Northern Cape.

  • funebris Kostylev

  • Quartinia funebris Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 101, 103, female, male –“Imam-Baba (Transkaspien)” (St. Petersburg).

  • Distribution: Turkmenistan.

  • galataea (Richards), new combination

  • Quartinioides galataea Richards, 1962, Revision. Study Masarid Wasps: 173 (key), 188, female –“Cape Province: Worcester” (London).

  • Distribution: South Africa: Western Cape.

  • goleana Richards

  • Quartinia goleana Richards, 1962, Revision. Study Masarid Wasps: 133, 134 (key), 137, female –“Algeria: 80 Km. nord d'El Goléa” (Paris).

  • Distribution: Algeria.

  • guichardi Richards

  • Quartinia guichardi Richards, 1969, J. Nat. Hist. 3: 80, female, male –“Canary Is., Gran Canaria, Maspalomas” (holotype male, London).

  • Distribution: Spain: Canary Islands: Gran Canaria.

  • haemorrhoa Gusenleitner

  • Quartinia haemorrhoa Gusenleitner, 1997, Linz. Biol. Beitr. 29(1): 137, 144, female –“Israel: Dead Sea, En Zeelim, 5 km N Massada, 31°23;pr/35°20;pr” (coll. M. Hauser).

  • Distribution: Israel.

  • halicticeps Giordani Soika

  • Quartinia halicticeps Giordani Soika, 1939, Bull. Soc. Fouad 1er Entomol. 1939: 9 (key), 12, figs. 2–4, female –“Egitto: Gabal Elba (Wadi Abrag)” (Venice, coll. Ministero de Agricoltura).

  • Distribution: Egypt.

  • hecuba (Richards), new combination

  • Quartinioides hecuba Richards, 1962, Revision. Study Masarid Wasps: 174 (key), 191, female, male –“Cape Province: Matjesfontein” (holotype male, London); also from Calvinia, and Kwazulu-Natal (“probably this species”).

  • Distribution: South Africa: Northern Cape, Western Cape, Kwazulu-Natal.

  • helena (Richards), new combination

  • Quartinioides helena Richards, 1962, Revision. Study Masarid Wasps: 173 (key), 187, female –“Cape Province: Cape Town” (Washington).

  • Distribution: South Africa: Western Cape.

  • helichrysi (Richards), new combination

  • Quartinioides helichrysi Richards, 1962, Revision. Study Masarid Wasps: 175 (key), 198, female –“Basutoland: Mamalapi Mts.” (London).

  • Distribution: Lesotho.

  • hetaira (Richards), new combination

  • Quartinioides hetaira Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 200, female, male –“Cape Province: Cape Town” (holotype male, Washington).

  • Distribution: South Africa: Western Cape.

  • hypatia Richards

  • Quartinia hypatia Richards, 1962, Revision. Study Masarid Wasps: 130, 132 (key), 160, male –“Cape Province: Calvinia” (London).

  • Distribution: South Africa: Northern Cape.

  • indica Cameron

  • Juartinia[!] indica Cameron, 1904, Z. Hym. Dipt. 4: 89, male –“Deesa, Western India” (London).

  • Quartinia maculifrons Dover, 1925 (1924), J. Proc. Asiat. Soc. Bengal (n. ser.) 20(6): 290. Unavailable under Art. 11.5 of the International Code of Zoological Nomenclature.

  • Distribution: India: Gujarat (Deesa).

  • interrupta Turner

  • Quartinia interrupta Turner, 1939, Ann. Transvaal Mus. 20: 2, female –“Willowmore, Cape Province” (lectotype, Pretoria).

  • Distribution: Namibia; South Africa: Western Cape, Eastern Cape.

  • iphigenia (Richards), new combination

  • Quartinioides iphigenia Richards, 1962, Revision. Study Masarid Wasps: 173, 174, 175 (key), 192, male –“S.W. Africa: Aus” (London).

  • Distribution: Namibia.

  • jocasta Richards

  • Quartinia jocasta Richards, 1962, Revision. Study Masarid Wasps: 130 (key), 160, female –“Cape Province: Bloukrans, near Calvinia” (London); also from Calvinia.

  • Distribution: South Africa: Northern Cape.

  • laeta von Schulthess

  • Quartinia capensis var. læta von Schulthess, 1935, Ann. Mag. Nat. Hist. (10) 16: 386 (key), 389, male, female –“Cape Province, Aus” (holotype male, London).

  • Distribution: Namibia.

  • latona (Richards), new combination

  • Quartinioides latona Richards, 1962, Revision. Study Masarid Wasps: 175, 177 (key), 207, female –“Cape Province: Prince Albert Road” (London).

  • Distribution: South Africa: Western Cape.

  • lesnei Benoist

  • Quartinia Lesnei Benoist, 1929, Bull. Soc. Sci. Nat. Maroc 9: 92, fig. 1, female –“Algérie: Tadmit” (Paris).

  • Distribution: Algeria.

  • libanica Richards

  • Quartinia libanica Richards, 1964 (1962), Boll. Mus. Civ. Stor. Nat. 15: 83, male, female –“N. Lebanon: Becharré, 5000 ft.” (holotype male, Venice); also from elsewhere in Lebanon, and Cyprus.

  • Distribution: Cyprus; Lebanon.

  • maerens von Schulthess

  • Quartinia capensis var. mærens von Schulthess, 1935, Ann. Mag. Nat. Hist. (10) 16: 386 (key), 389, female –“Cape Province, Worcester” (London); also from Matjesfontein.

  • Distribution: South Africa: Western Cape.

  • major Kohl

  • Quartinia major Kohl, 1898, Termesz. Füzetek 21: 363, pl. XV figs. 8, 21, female –“Oran … Auf Calendula” (Vienna).

  • ?Quartinia kohli Dusmet, 1917, Mem. R. Soc. Españ. Hist. Nat. 8., Mem. 9: 383, male –“Imi-n-Tanut” (Madrid).

  • Distribution: Morocco; Algeria.

  • matabele Turner

  • Quartinia matabele Turner, 1939, Ann. Transvaal Mus. 20: 3, male, female –“Bulawayo, S. Rhodesia” (Pretoria).

  • Distribution: Zimbabwe.

  • media von Schulthess

  • Quartinia media von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 506, female –“Cape Province, Montagu” (London).

  • Distribution: South Africa: Western Cape.

  • medusa Richards

  • Quartinia medusa Richards, 1962, Revision. Study Masarid Wasps: 135 (key), 159, figs. 152–153, female, male –“S.W. Africa: Aus” (holotype male, London).

  • Distribution: Namibia.

  • metallescens von Schulthess

  • Quartinia metallescens von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 505, female –“Cape Province, Queenstown, 3500 feet” (London).

  • Distribution: South Africa: Eastern Cape; Lesotho.

  • metope (Richards), new combination

  • Quartinioides metope Richards, 1962, Revision. Study Masarid Wasps: 175, 178 (key), 207, female –“S.W. Africa: Aus” (London).

  • Distribution: Namibia.

  • minima von Schulthess

  • Quartinia minima von Schulthess, 1932, Ann. Mag. Nat. Hist. (10) 10: 528, female –“Aus” (London).

  • Distribution: Namibia.

  • minuscula (Turner), new combination

  • Quartiniella minuscula Turner, 1939, Ann. Transvaal Mus. 20: 3, female –“Willowmore, Cape Province” (Pretoria).

  • Distribution: South Africa: Eastern Cape.

  • mochii Giordani Soika

  • Quartinia Mochii Giordani Soika, 1939, Bull. Soc. Fouad 1er Entomol. 1939: 9 (key), 10, fig. 1, male –“Egitto: Abu Rawash” (Venice, coll. Mochi).

  • Distribution: Egypt.

  • mongolica (Morawitz)

  • Celonites? mongolicus Morawitz, 1889, Horae Soc. Entomol. Ross. 23: 166, female, male –“Mongolia merid. Dshin-Tasy” (St. Petersburg).

  • Distribution: Mongolia.

  • multipicta (Richards), new combination

  • Quartinioides multipicta Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 202, female, male –“Cape Province: Worcester” (holotype male, London); also from elsewhere in Western Cape.

  • Distribution: South Africa: Western Cape.

  • nilotica Fischer

  • Quartinia nilotica Fischer, 1964, Ann. Naturhist. Mus. Wien 67: 611, fig. 1., female –“Khor Musa Pascha S. v. Wadi-Halfa” (Vienna).

  • Distribution: Egypt.

  • niveopicta von Schulthess

  • Quartinia niveopicta von Schulthess, 1930, Ann. Mag. Nat. Hist. (10) 5: 327 (key), 329, female –“Cape Province, Matjestfontein” (London).

  • Distribution: South Africa: Western Cape, Eastern Cape.

  • nubiana Richards

  • Quartinia nubiana Richards, 1962, Revision. Study Masarid Wasps: 133, 134 (key), 138, female –“Egypt: Nubian Desert, Korosko to Abu Hamed” (London); also from elsewhere in Egypt.

  • Distribution: Tunisia; Libya; Egypt; Sudan; Saudi Arabia; United Arab Emirates.

  • ochraceopicta von Schulthess

  • Quartinia ochraceopicta von Schulthess, 1932, Ann. Mag. Nat. Hist. (10) 10: 527, female –“Aus” (London).

  • Distribution: Namibia.

  • orientalis Gusenleitner

  • ?“Quartinia haemorrhoa (n. sp.) frontalis n. ssp.” Blüthgen, 1961, Beitr. Naturk. Forsch. S.W.-Deutschl. 19(3): 281, male –“Gondahar.” Nomen nudum.

  • Quartinia orientalis Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 56, fig. 1b, male, female –“Türkei Ürgüp” (holotype male, Linz); also from Afghanistan.

  • Distribution: Turkey; Afghanistan.

  • paradoxa Brauns

  • Quartinia paradoxa Brauns, 1905, Ann. Mus. Natl. Hung. 3: 233, male –“Terra Capensis: Willowmore” (Pretoria).

  • Distribution: South Africa: Eastern Cape.

  • parcepunctata Richards

  • Quartinia parcepunctata Richards, 1962, Revision. Study Masarid Wasps: 133 (key), 167, female, male –“Cape Town” (holotype female, Washington).

  • Distribution: South Africa: Western Cape.

  • parvula Dusmet

  • Quartinia parvula Dusmet, 1909, Act. Mem. Primer Congr. Nat. Españ., Zaragoza, 1908: 183, male, female (Madrid).

  • Distribution: Portugal; Spain.

  • perone Richards

  • Quartinia perone Richards, 1962, Revision. Study Masarid Wasps: 133, 134 (key), 166 –“Cape Province: Worcester” (London).

  • Distribution: South Africa: Western Cape.

  • persephone Richards

  • Quartinia persephone Richards, 1962, Revision. Study Masarid Wasps: 133 (key), 165, female –“Cape Province: Calvinia” (London).

  • Distribution: South Africa: Northern Cape.

  • philomela (Richards), new combination

  • Quartinioides philomela Richards, 1962, Revision. Study Masarid Wasps: 173, 174 (key), 193, female –“Cape Province: Nieuwoudtville” (London).

  • Distribution: South Africa: Northern Cape.

  • phoebe (Richards), new combination

  • Quartinioides phoebe Richards, 1962, Revision. Study Masarid Wasps: 172 (key), 181, male –“Cape Province: Cape Town” (Washington).

  • Distribution: South Africa: Western Cape.

  • pluto Richards

  • Quartinia pluto Richards, 1962, Revision. Study Masarid Wasps: 133, 134 (key), 166, female –“Cape Province: Worcester” (London).

  • Distribution: South Africa: Western Cape.

  • poecila von Schulthess

  • Quartinia poecila von Schulthess, 1930, Ann. Mag. Nat. Hist. (10) 5: 327, fig. 2, female, male –“Cape Province, Swakopmund” (lectotype male, London).

  • Distribution: Namibia.

  • popovi Richards

  • Quartinia popovi Gussakovskii, 1935, Trudy Tadzik. Sect. Acad. Sci. U.S.S.R. (Zool. Parasitol.) 5: 466, fig. 23. Nomen nudum.

  • Quartinia popovi Richards, 1962, Revision. Study Masarid Wasps: 130 (key), 144, female –“District of Kondara” (Moscow).

  • Distribution: Tajikistan.

  • propinqua von Schulthess

  • Quartinia propinqua von Schulthess, 1932, Ann. Mag. Nat. Hist. (10) 10: 526, figs. 2–4, male, female –“Aus” (lectotype female, London).

  • Distribution: Namibia; South Africa: Western Cape.

  • proserpina Richards

  • Quartinia proserpina Richards, 1962, Revision. Study Masarid Wasps: 133 (key), 161, female –“Cape Province: Nieuwoudtville” (London).

  • Distribution: South Africa: Northern Cape.

  • punctulata von Schulthess

  • Quartinia punctulata von Schulthess, 1930, Ann. Mag. Nat. Hist. (10) 5: 327 (key), 329, female –“Cape Province, Matjesfontein” (London).

  • Quartinia punctulatum[!] Turner, 1939, Ann. Transvaal Mus. 20: 1.

  • Distribution: South Africa: Western Cape.

  • pusilla Kostylev

  • Quartinia puslilla[!]; Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 101. Incorrect original spelling.

  • Quartinia pusilla Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 103 (key), 104, female –“Krasnovodsk” (St. Petersburg).

  • Distribution: Turkmenistan.

  • senecionis (Richards), new combination

  • Quartinioides senecionis Richards, 1962, Revision. Study Masarid Wasps: 176 (key), 205, female, male –“Orange Free State: Thaba Nchu” (holotype male, London); also from Lesotho.

  • Distribution: Lesotho; South Africa: Free State.

  • separata Gusenleitner

  • Quartinia separata Gusenleitner, 1997, Linz. Biol. Beitr. 29(1): 137, 146, female, male –“Syria west, 50 km S Homs” (holotype female, BOL); also from elsewhere in Syria; Turkey.

  • Distribution: Syria; Turkey.

  • shestakovi Kostylev

  • Quartinia shestakovi Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 101, 102 (key), 105, fig. 8, female, male –“Tartugaj (Syr-Darjja-Geb.)” (St. Petersburg); also from elsewhere in Uzbekistan.

  • Quartinia schestakovi[!]; Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 105. Incorrect original spelling.

  • Distribution: Uzbekistan; Kyrgyzstan.

  • signata von Schulthess

  • ?Quartinia variegata “Brauns” Bradley, 1922, Univ. Calif. Publ. Entomol. 1: 401. Nomen nudum.

  • Quartinia signata, Brauns, in litt.” von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 507, male, female –“Cape Province, Willowmore” (lectotype male, Zürich).

  • Distribution: South Africa: Eastern Cape.

  • signatifrons Turner

  • Quartinia signata var. signatifrons Turner, 1939, Ann. Transvaal Mus. 20: 2, male, female (Pretoria).

  • Distribution: South Africa: Western Cape, Eastern Cape.

  • soikai Richards

  • Quartinia soikai Richards, 1964 (1962), Boll. Mus. Civ. Stor. Nat. 15: 84, female –“Turkey: Steppe hills 70 m. S. of Ankara” (Venice).

  • Quartinia soika[!]; Gusenleitner, 1985, Nachricht. Bayer. Entomol. 34(4): 110.

  • Distribution: Iran; Turkey; Greece.

  • striata (Richards), new combination

  • Quartiniella striata Richards, 1962, Revision. Study Masarid Wasps: 208 (key), 211, female –“Cape Province: Calvinia” (London).

  • Distribution: South Africa: Northern Cape.

  • syriaca nitens Gusenleitner

  • Quartinia syriaca nitens Gusenleitner, 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 57, male –“Iran, Fars, Monti steppici 5 km N di Persepolis” (Venice).

  • Distribution: Iran.

  • syriaca syriaca Richards

  • Quartinia syriaca Richards, 1964 (1962), Boll. Mus. Civ. Stor. Nat. 15: 86, female, male –“Syria: near Damascus, Ibn Nefis” (holotype female, Venice); also from other localities in Syria, and Lebanon.

  • Distribution: Lebanon; Syria.

  • tarsata (Richards), new combination

  • Quartinioides tarsata Richards, 1962, Revision. Study Masarid Wasps: 173 (key), 185, female, male –“Cape Province: Grahamstown” (holotype female, London).

  • Distribution: South Africa: Eastern Cape.

  • tenerifina Richards

  • Quartinia tenerifina Richards, 1969, J. Nat. Hist. 3: 80, fig. 2, female, male –“Canary Is., Tenerife, Los Christianos” (holotype male, London).

  • Distribution: Spain: Canary Islands: Tenerife.

  • thebaica du Buysson

  • Quartinia thebaica du Buysson, 1902, Bull. Soc. Entomol. France 1902: 141, female –“Égypte: Louqsor” (coll. Pic).

  • Quartinia thebica[!]; Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 102.

  • Quartinia ochracea Giordani Soika, 1939, Bull. Soc. Fouad 1er Entomol. 1939: 9 (key), 14, female –“Egitto: Wadi Digla” (Venice, coll. Mochi).

  • Quartinia thabaica[!]; Gusenleitner, 1977, Linz. Biol. Beitr. 9: 152.

  • Distribution: Morocco; Algeria; Libya; Egypt.

  • titania (Richards), new combination

  • Quartinioides titania Richards, 1962, Revision. Study Masarid Wasps: 174 (key), 193, female, male –“Cape Province: Cape Town” (holotype male, Washington).

  • Distribution: South Africa: Western Cape.

  • tricolorata Giordani Soika

  • Quartinia thebaica du Buysson; Giordani Soika, 1939, Bull. Soc. Fouad 1er Entomol. 39: 13. Misidentification.

  • Quartinia tricolorata Giordani Soika, 1954, Boll. Mus. Civ. Stor. Nat. Venezia 7: 22, female –“Egitto: Gebel Asfar” (lectotype Venice); also from Fayum.

  • Distribution: Egypt.

  • tripolitana sinaitica Richards

  • Quartinia tripolitana sinaitica Richards, 1964 (1962), Boll. Mus. Civ. Stor. Nat. 15: 83, male, female –“United Arab Republic, Sinai: Kun Mitla” (Venice).

  • Distribution: Egypt: Sinai.

  • tripolitana tripolitana Richards

  • Quartinia tripolitana Richards, 1962, Revision. Study Masarid Wasps: 134 (key), 140, male, female –“Tripolitania: Tripoli” (holotype male, London); also from elsewhere in Libya.

  • Distribution: Libya; Egypt.

  • tuareg Giordani Soika

  • Quartinia tuareg Giordani Soika, 1954, Boll. Mus. Civ. Stor. Nat. Venezia 7: 21, male, female –“Guelta Idjef Melène” (lectotype female, Venice); also from elsewhere in Algeria.

  • Distribution: Algeria; Egypt.

  • turneri (von Schulthess)

  • Quartiniella turneri von Schulthess, 1932, Ann. Mag. Nat. Hist. (10) 10: 528, fig. 5, male –“Aus” (London).

  • Distribution: Namibia.

  • uzbeka Kostylev

  • Quartinia uzbeka Kostylev, 1935, Arch. Mus. Zool. Univ. Moscou 2: 101, 103 (key), 106, male –“Aman-Katan, Umgesb. v. Samarkand, 1.500 m. üb. M.” (St. Petersburg).

  • Distribution: Uzbekistan.

  • vagepunctata von Schulthess

  • Quartinia vagepunctata von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 507, female –“Cape Province, Little Karroo, 38 m. E. of Ceres” (London).

  • Distribution: South Africa: Northern Cape, Western Cape.

  • waterstoni (von Schulthess)

  • Quartiniella waterstoni von Schulthess, 1929, Ann. Mag. Nat. Hist. (10) 3: 504 (key), 510, figs. 4–5, female, male –“Cape Province, Little Karroo, 38 m. E. of Ceres” (holotype female, London).

  • Distribution: South Africa: Northern Cape, Western Cape, Kwazulu-Natal.

  • Genus Trimeria de Saussure

  • Trimeria de Saussure, 1854, Ét. Fam. Vesp. 3: pl. IV fig. 2; 1855, Ét. Fam. Vesp. 3: 81, genus. Type species: Erynnis americana de Saussure, 1853, by monotypy.

  • Erynnis de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xix, genus (2 species), non Erynnis Schrank, 1801 (Lepidoptera). Type species: Erynnis americana de Saussure, 1853, by subsequent designation of de Saussure, 1855, Ét. Fam. Vesp. 3: 92.

  • Eurynnis[!] Dalla Torre, 1894, Cat. Hym. 9: 4.

  • Microtrimeria Bequaert, 1928, Psyche 35: 191, genus: 191. Type species: Microtrimeria cockerelli Bequaert, 1928, by monotypy.

  • Willinkia Richards, 1962, Revision. Study Masarid Wasps: 119, subgenus of Trimeria de Saussure. Type species:Trimeria buyssoni Brèthes, 1904 (as represented in coll. B. M.),” by original designation.

  • americana (de Saussure)

  • Masaris vespiformis Fabricius; Romand, 1851, Ann. Soc. Entomol. France (2) 9, Bull.: lii (female not male). Misidentification.

  • Erynnis americana de Saussure, 1853, Ann. Soc. Entomol. France (3) 1, Bull. II: xx, female –“d'Amérique” (Paris).

  • Distribution: Brazil: Goiás,* Distrito Federal,* São Paulo,* Paraná.*

  • atacama (Fritz), new combination

  • Microtrimeria atacama Fritz, 1968, Rev. Chil. Entomol. 6: 69, female, male –“Estación Cachiyuyo, provincia de Atacama, Chile” (holotype female, Santiago).

  • Distribution: Chile: Atacama.

  • bequaerti Willink

  • Trimeria bequaerti Willink, 1951, Rev. Soc. Cient. Argentina 15: 78, figs. 1–6, female –“Amaicha del Valle, provincia de Tucumán (2.000 ms, s.n.m.)” (Tucumán); also from Salta.

  • Distribution: Bolivia; Paraguay: Boquerón; Argentina: Salta, Tucumán.

  • cockerelli (Bequaert), new combination

  • Microtrimeria cockerelli Bequaert, 1928, Psyche 35: 191, 192, female –“PERU: Payta” (Cambridge).

  • Distribution: Peru: Piura.

  • howardi Bertoni

  • Trimeria howardi Bertoni, 1911, An. Mus. Nac. Buenos Aires (3) 22: 104, female, male –“Puerto Bertoni” (San Lorenzo).

  • Distribution: Brazil*: Minas Gerais, São Paulo, Rio Grande do Sul; Paraguay: Alto Paraná; Argentina: Tucumán, Santa Fé, Misiones.

  • joergenseni Schrottky

  • Trimeria joergenseni Schrottky, 1909, Rev. Mus. La Plata 16: 137, female –“Argentina (Mendoza)” (destroyed).

  • Distribution: Argentina: Mendoza, San Luis,* Santa Fé,* Rio Negro.

  • monrosi Willink

  • Trimeria monrosi Willink, 1959, Neotropica 5: 2, female –“Tucumán, Las Cejas” (Tucumán).

  • Distribution: Bolivia: Santa Cruz; Argentina: Tucumán, Santiago del Estero,* Santa Fé.

  • neotropica (Mocsáry)

  • Jugurtia neotropica Mocsáry, 1906, Ann. Hist.-Nat. Mus. Hung. 4: 197, male –“Paraguay: Asuncion” (Budapest).

  • Distribution: Brazil*: Paraná; Paraguay: Central; Argentina*: Misiones.

  • rhachiphora (Schletterer)

  • Odynerus rhachiphorus Schletterer, 1891, Entomol. Nachr. 17: 85, male –“Chile” (Vienna).

  • Trimeria buyssoni Brèthes, 1903, An. Mus. Nac. Buenos Aires (3) 9(2): 371, figs. 1–2, 372, fig. 3, female –“Argentina: Catamarca” (Buenos Aires); also described from Córdoba? and Paraguay.

  • Trimeria rachiphorus[!]; Willink and Roig Alsina, 1993, J. Hym. Res. 2: 303.

  • Distribution: Paraguay: Boquerón, Itapuá; Uruguay*: Montevideo; Argentina: Jujuy, Salta, Tucumán, Catamarca, Santiago del Estero, Formosa, Santa Fé, Neuquén.

  • Acknowledgments

    I thank Mike Engel, Fred Gess, Josef Gusenleitner, Norm Johnson, Lynn Kimsey, and Rob Longair for critical comments, and Vladimir Ovtsharenko for help with Russian localities.

    REFERENCES

    1.

    J. M. Carpenter 1981 (1982). The phylogenetic relationships and natural classification of the Vespoidea (Hymenoptera). Syst. Entomol 7:11–38. Google Scholar

    2.

    J. M. Carpenter 1989 (1988). The phylogenetic system of the Gayellini (Hymenoptera: Vespidae; Masarinae). Psyche 95:211–241. Google Scholar

    3.

    J. M. Carpenter 1993. Biogeographic patterns in the Vespidae (Hymenoptera): two views of Africa and South America. In P. Goldblatt (ed.), Biological Relationships between Africa and South America:. 139–155. New Haven, CT: Yale Univ. Press. Google Scholar

    4.

    J. M. Carpenter 1997 (1996). Generic classification of the Australian pollen wasps (Hymenoptera: Vespidae; Masarinae). J. Kansas Entomol. Soc 69(4) suppl.:384–400. Google Scholar

    5.

    J. S. Farris 1979. The information content of the phylogenetic system. Syst. Zool 28:483–519. Google Scholar

    6.

    F. W. Gess 1997. Contributions to knowledge of Ceramius Latreille, Celonites Latreille Jugurtia Saussure and Masarina Richards (Hymenoptera: Vespidae: Masarinae) in South Africa. J. Hym. Res 6:36–74. Google Scholar

    7.

    F. W. Gess 1998. Priscomasaris namibiensis Gess, a new genus and species of Masarinae (Hymenoptera: Vespidae) from Namibia, southern Africa, with a discussion of its position within the subfamily. J. Hym. Res 7:296–304. Google Scholar

    8.

    F. W. Gess and S. K. Gess . 1988. A contribution to the knowledge of the taxonomy and ethology of the genus Masarina Richards (Hymenoptera: Masaridae). Ann. Cape Prov. Mus. (Nat. Hist.) 16:351–362. Google Scholar

    9.

    S. K. Gess 1996. The pollen wasps: ecology and natural history of the Masarinae. Cambridge, MA: Harvard Univ. Press. Google Scholar

    10.

    A. Giordani Soika 1974. Revisione della sottofamiligia Gayellinae (Hym. vesp.). Boll. Mus. Civ. Stor. Nat. Venezia 25:87–106. Google Scholar

    11.

    O. W. Richards 1962. A revisional study of the masarid wasps (Hymenoptera, Vespoidea). London: British Museum (Natural History). Google Scholar

    12.

    R. R. Snelling 1986. The taxonomy and nomenclature of some Australian paragiine wasps (Hymenoptera, Masaridae). Contrib. Sci 378:1–19. Google Scholar

    13.

    Jvan der Vecht and J. M. Carpenter . 1990. A catalog of the genera of the Vespidae (Hymenoptera). Zool. Verh. (Leiden) 260:1–62. Google Scholar

    Appendices

    INDEX OF MASARINAE

    Genera, species, and subspecies are indexed, including emendations, misspellings, and lapsi calami, etc. The generic name given in parentheses following a species-group entry is the name used in the original combination. The name of the genus that now includes the species-group taxon is then listed below. If the species-group taxon is now considered a synonym, the valid name is listed following the current genus name. Similarly, for emendations, misspellings, and lapsi, the correct spelling follows. If the species-group taxon is considered a subspecies, this is indicated by the abbreviation “ssp.” following the name of the nominate species.

    • abbreviata (Vespa)

    •  Celonites abbreviatus, 8

    • abbreviatus tauricus (Celonites)

    •  Celonites abbreviatus ssp., 8

    • abbreviatus var. engadinensis (Celonites)

    •  Celonites abbreviatus, 8

    • abbreviatus var. hungaricus (Celonites)

    •  Celonites abbreviatus, 8

    • afer (Celonites)

    •  Celonites 9

    • affinis (Quartinia)

    •  Quartinia 22

    • albifrons (Pseudomasaris)

    •  Pseudomasaris maculifrons 21

    • albopicta (Quartinioides)

    •  Quartinia 22

    • alcestis (Quartinia)

    •  Quartinia 22

    • alfkeni (Masaris)

    •  Jugurtia 17

    • algerica (Masariella?)

    •  Jugurtia 17

    • Alpha

    •  Paragia 6

    • americana (Erynnis)

    •  Trimeria 29

    • Ammoparagia

    •  Metaparagia 5

    • andrei (Celonites)

    •  Celonites 9

    • andromeda (Quartinioides)

    •  Quartinia 22

    • angulata (Riekia)

    •  Metaparagia 5

    • antennata (Quartinia)

    •  Quartinia 23

    • antigone (Quartinioides)

    •  Quartinia 23

    • apiformis (Masaris)

    •  Celonites abbreviatus 8

    • araucana (Gayella)

    •  Gayella 4

    • araxana (Quartinia)

    •  Quartinia 23

    • arsinoe (Quartinioides)

    •  Quartinia 23

    • artemis (Quartinia)

    •  Quartinia 23

    • atacama (Microtrimeria)

    •  Trimeria 30

    • atra (Quartinia)

    •  Quartinia 23

    • atrata (Quartinia)

    •  Quartinia 23

    • aucta (Liris)

    •  Ceramius auctus 14

    • australis (Paragia)

    •  Paragia 6

    • australis borealis (Paragia)

    •  Paragia australis ssp. 6

    • bariscapus[!] (Pseudomasaris)

    •  Pseudomasaris basirufus 21

    • bariscipus (Pseudomasaris)

    •  Pseudomasaris basirufus 21

    • basuto (Quartinioides)

    •  Quartinia 23

    • beaumonti (Paraceramius)

    •  Ceramius 14

    • bequaerti (Trimeria)

    •  Trimeria 30

    • bergenwahliae (Celonites)

    •  Celonites 9

    • Beta

    •  Paragia 6

    • beyeri (Ceramius)

    •  Ceramius 14

    • bicolor (Paragia)

    •  Paragia 6

    • bicolor (Philanthus)

    •  Ceramius 14

    • bidens (Paragia)

    •  Paragia odyneroides 7

    • bischoffi (Ceramius)

    •  Ceramius 14

    • biskrensis (Jugurtia)

    •  Jugurtia 17

    • borreriae (Rolandia)

    •  Metaparagia 5

    • brasiliensis (Paramasaris)

    •  Paramasaris 5

    • braunsi (Ceramiellus)

    •  Jugurtia 17

    • braunsi (Ceramius)

    •  Ceramius 14

    • braunsiella (Masaris)

    •  Jugurtia 17

    • brevitarsis (Ceramius)

    •  Ceramius 14

    • breyeri (Quartinia)

    •  Quartinia 23

    • bureschi (Ceramius)

    •  Ceramius 14

    • bureschi lycaonius (Ceramius)

    •  Ceramius bureschi 14

    • buyssoni (Trimeria)

    •  Trimeria rhachiphora 30

    • caffer (Ceramius)

    •  Ceramius 14

    • calcarata (Jugurtia)

    •  Jugurtia 17

    • calida (Paragia)

    •  Paragia 6

    • canariensis (Quartinia)

    •  Quartinia 23

    • candida (Quartinia)

    •  Quartinia 23

    • capensis (Celonites)

    •  Celonites 9

    • capensis (Ceramius)

    •  Ceramius 14

    • capensis (Quartinia)

    •  Quartinia 23

    • capensis var. laeta (Quartinia)

    •  Quartinia laeta 25

    • capensis var. mærens (Quartinia)

    •  Quartinia maerens 26

    • capicola (Ceramius)

    •  Ceramius 14

    • carli (Masaris)

    •  Masaris 20

    • caucasicus (Ceramius)

    •  Ceramius 14

    • cazieri (Pseudomasaris)

    •  Pseudomasaris 21

    • Celonites 8

    • Celonitinae

    •  Masarinae 4

    • Ceramias[!]

    •  Ceramius 13

    • Ceramiellus

    •  Jugurtia 17

    • Ceramioides

    •  Ceramius 13

    • Ceramiopsis 13

    • Ceramius 13

    • Ceramus[!]

    •  Ceramius 13

    • cerceriformis (Ceramius)

    •  Ceramius 14

    • cerceroides (Gayella)

    •  Gayella patagonica 4

    • ceres (Masarina)

    •  Masarina 19

    • ceres (Quartinioides)

    •  Quartinia 23

    • Chelonites[!]

    •  Celonites 8

    • chlorotica (Jugurtia)

    •  Quartinia 23

    • cincta (Quartinia)

    •  Quartinia 24

    • clarus (Celonites)

    •  Celonites 9

    • clypeatus (Celonites)

    •  Celonites 9

    • clypeatus (Ceramius)

    •  Ceramius 15

    • cockerelli (Microtrimeria)

    •  Trimeria 30

    • codoni (Jugurtia)

    •  Jugurtia 17

    • Coelonites[!]

    •  Celonites 8

    • communis (Masarina)

    •  Masarina familiaris 19

    • concinna (Paragia)

    •  Paragia venusta 8

    • confluens (Paragia)

    •  Metaparagia 5

    • confusa (Jugurtia)

    •  Jugurtia 17

    • coquilletti (Pseudomasaris)

    •  Pseudomasaris 21

    • Cotyledon

    •  Pseudomasaris 21

    • crenulatus (Celonites)

    •  Celonites 9

    • cressida (Quartinioides)

    •  Quartinia 24

    • cretica (Quartinia)

    •  Quartinia 24

    • cupreus (Paramasaris)

    •  Paramasaris 5

    • Cygnaea

    •  Paragia 6

    • cyllene (Quartinioides)

    •  Quartinia 24

    • cynara (Quartinioides)

    •  Quartinia 24

    • cyprius (Celonites)

    •  Celonites 9

    • cyprius smyrnensis (Celonites)

    •  Celonites cyprius ssp. 9

    • damarinus (Ceramius)

    •  Ceramius 15

    • davidi (Celonites)

    •  Celonites 9

    • deceptor (Paragia)

    •  Paragia 6

    • deceptrix (Paragia)

    •  Paragia deceptor 6

    • decipiens (Paragia)

    •  Paragia 7

    • decipiens aliciae (Paragia)

    •  Paragia decipiens ssp. 6

    • diana (Quartinioides)

    •  Quartinia 24

    • dilecta (Quartinia)

    •  Quartinia 24

    • discrepans (Masaris)

    •  Jugurtia 17

    • discretus (Celonites)

    •  Celonites 9

    • dispar (Celonites)

    •  Jugurtia 18

    • doddi (Metaparagia)

    •  Metaparagia 6

    • dryope (Quartinioides)

    •  Quartinia 24

    • dubia (Chrysis)

    •  Celonites abbreviatus 8

    • duplicata (Jugurtia)

    •  Jugurtia 18

    • edwardsii (Masaris)

    •  Pseudomasaris 21

    • elissa (Quartinioides)

    •  Quartinia 24

    • eremobia (Quartinia)

    •  Quartinia 24

    • Erynnis

    •  Trimeria 29

    • escalerae (Jugurtia)

    •  Jugurtia 18

    • Eucelonites

    •  Celonites 8

    • Euceramius

    •  Ceramius 13

    • eumenoides (Gayella)

    •  Gayella 4

    • eurycara (Jugurtia)

    •  Jugurtia 18

    • eurydice (Quartinioides)

    •  Quartinia 24

    • Eurynnis[!]

    •  Trimeria 29

    • excellens (Paragia)

    •  Paragia sobrina 8

    • familiaris (Masarina)

    •  Masarina 19

    • fischeri (Celonites)

    •  Celonites 9

    • flava (Quartiniella)

    •  Quartinia 24

    • flavolineatus (Zethoides)

    •  Paramasaris fuscipennis 5

    • fonscolombei (Ceramius)

    •  Ceramius 15

    • fonscolombei oraniensis (Ceramius)

    •  Ceramius fonscolombei ssp. 15

    • foveolatus (Celonites)

    •  Celonites 9

    • foveolatus nigrior (Celonites)

    •  Celonites foveolatus ssp. 9

    • fumipennis (Ceramius)

    •  Ceramius linearis 15

    • funebris (Quartinia)

    •  Quartinia 24

    • fuscipennis (Paramasaris)

    •  Paramasaris 5

    • galataea (Quartinioides)

    •  Quartinia 24

    • gariepensis (Celonites)

    •  Celonites 9

    • Gayella 4

    • Gayellini 4

    • Gazella[!]

    •  Gayella 4

    • generosa (Paragia)

    •  Paragia 7

    • gessi (Ceramius)

    •  Ceramius 15

    • gestroi (Ceramiopsis)

    •  Ceramiopsis 13

    • Gnatho

    •  Ceramius 13

    • goleana (Quartinia)

    •  Quartinia 24

    • guichardi (Celonites)

    •  Celonites 10

    • guichardi (Quartinia)

    •  Quartinia 25

    • gussakovskii (Masaris)

    •  Masaris 20

    • haemorrhoa (Quartinia)

    •  Quartinia 25

    • haemorrhoa frontalis (Quartinia)

    •  ?Quartinia orientalis 27

    • halicticeps (Quartinia)

    •  Quartinia 25

    • hamanni (Celonites)

    •  Celonites 10

    • hammani[!] (Celonites)

    •  Celonites hamanni 10

    • hecuba (Quartinioides)

    •  Quartinia 25

    • helena (Quartinioides)

    •  Quartinia 25

    • helichrysi (Quartinioides)

    •  Quartinia 25

    • hellenicus (Celonites)

    •  Celonites 10

    • hessei (Ceramius)

    •  Ceramius nigripennis 16

    • hetaira (Quartinioides)

    •  Quartinia 25

    • hirsuta (Paragia)

    •  Paragia 7

    • hispanicus (Ceramius)

    •  Ceramius 15

    • hoggarica (Jugurtia)

    •  Jugurtia jemenensis 18

    • Holopticus

    •  Pseudomasaris 21

    • houstoni (Rolandia)

    •  Metaparagia 6

    • howardi (Trimeria)

    •  Trimeria 30

    • hua (Ammoparagia)

    •  Metaparagia 6

    • humeralis (Celonites)

    •  Celonites 10

    • hyalinipennis (Masarina)

    •  Masarina 19

    • hylaeiformis (Ceramius)

    •  Masaris vespiformis 20

    • hypatia (Quartinia)

    •  Quartinia 25

    • hystrix (Celonites)

    •  Celonites 10

    • indica (Quartinia)

    •  Quartinia 25

    • interrupta (Quartinia)

    •  Quartinia 25

    • iphigenia (Quartinioides)

    •  Quartinia 25

    • irana (Jugurtia)

    •  Jugurtia 18

    • jacoti (Ceramius)

    •  Ceramius 15

    • jemenensis (Jugurtia)

    •  Jugurtia 18

    • jemenesis[!] (Jugurtia)

    •  Jugurtia jemenensis 18

    • jocasta (Quartinia)

    •  Quartinia 25

    • joergenseni (Trimeria)

    •  Trimeria 30

    • joussaumei[!] (Celonites)

    •  Celonites jousseaumei 10

    • jousseaumei (Celonites)

    •  Celonites 10

    • jousseaumei asrensis (Celonites)

    •  Celonites jousseaumei jousseaumei 10

    • jousseaumei senegalensis (Celonites)

    •  Celonites jousseaumei ssp. 10

    • Juartinia[!]

    •  Quartinia 22

    • Juguartia[!]

    •  Jugurtia 17

    • Jugurthia[!]

    •  Jugurtia 17

    • Jugurthia

    •  Jugurtia 17

    • Jugurtia 17

    • Jujurtha[!]

    •  Jugurtia 17

    • karooensis[!] (Ceramius)

    •  Ceramius bicolor 14

    • karrooensis (Ceramius)

    •  Ceramius bicolor 14

    • koeroegabensis (Jugurtia)

    •  Jugurtia 18

    • kohli (Quartinia)

    •  ? Quartinia major 26

    • kostylevi (Celonites)

    •  Celonites 10

    • kozlovi (Celonites)

    •  Celonites 10

    • laetus (Celonites)

    •  Celonites 10

    • latitarsis (Celonites)

    •  Celonites 10

    • latona (Quartinioides)

    •  Quartinia 25

    • lesnei (Quartinia)

    •  Quartinia 25

    • libanica (Quartinia)

    •  Quartinia 26

    • lichtensteinii (Gnatho)

    •  Ceramius 15

    • linearis (Ceramius)

    •  Ceramius 15

    • lobeliae (Celonites)

    •  Celonites 10

    • longicornis (Saryara)

    •  Masaris 20

    • longipilis (Celonites)

    •  Celonites 11

    • luispenai (Gayella)

    •  Gayella 4

    • lusitanicus (Ceramius)

    •  Ceramius 15

    • lusitanicus var. luteo---clypeata (Ceramius)

    •  Ceramius lusitanicus 15

    • macneilli (Pseudomasaris)

    •  Pseudomasaris 21

    • macrocephalus (Ceramius)

    •  Ceramius lichtensteinii 15

    • macswaini (Pseudomasaris)

    •  Pseudomasaris 21

    • maculata (Paragia)

    •  Metaparagia 6

    • maculifrons (Masaris)

    •  Pseudomasaris 21

    • maculifrons (Quartinia)

    •  Quartinia indica 25

    • magdalena (Paragia)

    •  Paragia 7

    • major (Quartinia)

    •  Quartinia 26

    • marginalis (Masaris)

    •  Pseudomasaris 21

    • maroccanus (Paraceramius)

    •  Ceramius 15

    • maroccanus rubripes (Ceramius)

    •  Ceramius rubripes 16

    • Masariella

    •  Jugurtia 17

    • Masarina 8 19

    • Masarini 5

    • Masaris 20

    • Massaris

    •  Masaris 20

    • matabele (Quartinia)

    •  Quartinia 26

    • mayeti (Celonites)

    •  Celonites 11

    • Mazaridinae

    •  Masarinae 4

    • Mazaris

    •  Masaris 20

    • media (Quartinia)

    •  Quartinia 26

    • medusa (Quartinia)

    •  Quartinia 26

    • metallescens (Quartinia)

    •  Quartinia 26

    • metanotalis (Ceramius)

    •  Ceramius 16

    • Metaparagia 5

    • Metazethoides

    •  Paramasaris 4

    • metope (Quartinioides)

    •  Quartinia 26

    • michaelseni (Celonites)

    •  Celonites 11

    • micheneri (Ceramius)

    •  Ceramius 16

    • micheneri (Pseudomasaris)

    •  Pseudomasaris 22

    • Microtrimeria

    •  Trimeria 29

    • mimetica (Paragia)

    •  Paragia 7

    • minima (Quartinia)

    •  Quartinia 26

    • minuscula (Quartiniella)

    •  Quartinia 26

    • mixta (Masarina)

    •  Masarina 19

    • mixtoides (Masarina)

    •  Masarina 19

    • mochii (Quartinia)

    •  Quartinia 26

    • modestus (Celonites)

    •  Celonites 11

    • modestus biinterruptus (Celonites)

    •  Celonites modestus ssp. 11

    • modestus bisinterruptus[!] (Celonites)

    •  Celonites modestus biinterruptus 11

    • mongolica (?Celonites)

    •  Quartinia 26

    • monocesta (Paragia)

    •  Paragia 7

    • monrosi (Trimeria)

    •  Trimeria 30

    • montanus (Celonites)

    •  Celonites 11

    • montanus (Ceramius)

    •  Ceramius 16

    • morosa (Paragia)

    •  Paragia 7

    • multipicta (Quartinioides)

    •  Quartinia 26

    • mutilloides (Gayella)

    •  Gayella 4

    • mutilloides nigerrima (Gayella)

    •  Gayella mutilloides 4

    • nadigorum (Jugurtia)

    •  Jugurtia 18

    • namaqua (Masarina)

    •  Masarina 19

    • namibiensis (Priscomasaris)

    •  Priscomasaris 5

    • nasuta (Paragia)

    •  Paragia 7

    • neotropica (Jugurtia)

    •  Trimeria 30

    • nigripennis (Ceramius)

    •  Ceramius 16

    • nilotica (Quartinia)

    •  Quartinia 26

    • niveopicta (Quartinia)

    •  Quartinia 26

    • nocatunga (Riekia)

    •  Metaparagia 6

    • nubiana (Quartinia)

    •  Quartinia 26

    • numida[!] (Jugurtia)

    •  Jugurtia oraniensis 18

    • numidica (Jugurtia)

    •  Jugurtia oraniensis 18

    • nursei (Celonites)

    •  Celonites 11

    • occidentalis (Masaris)

    •  Pseudomasaris 22

    • ochracea (Quartinia)

    •  Quartinia thebaica 28

    • ochraceopicta (Quartinia)

    •  Quartinia 27

    • octoannulata (Saryara)

    •  Celonites octoannulatus 11

    • octoannulatus hissaricus (Celonites)

    •  Celonites octoannulatus ssp. 11

    • odyneroides (Gayella)

    •  Gayella mutilloides 4

    • odyneroides (Paragia)

    •  Paragia 7

    • oligomera (Paragia)

    •  Paragia 7

    • oraniensis (Celonites)

    •  Jugurtia 18

    • orientalis (Quartinia)

    •  Quartinia 27

    • osseus (Celonites)

    •  Celonites 11

    • palaestinensis (Paraceramius)

    •  Ceramius 16

    • palestinensis[!] (Ceramius)

    •  Ceramius palaestinensis 16

    • Paracelonites

    •  Celonites 8

    • Paraceramius

    •  Ceramius 13

    • paradoxa (Quartinia)

    •  Quartinia 27

    • Paragayella

    •  Paramasaris 5

    • Paragia 6

    • Paragiella

    •  Paragia 6

    • Paragiina 5

    • paraguayensis (Ceramiopsis)

    •  Ceramiopsis 13

    • Paramasaris 4

    • parcepunctata (Quartinia)

    •  Quartinia 27

    • parvula (Masarina)

    •  Masarina 19

    • parvula (Quartinia)

    •  Quartinia 27

    • patagonica (Gayella)

    •  Gayella 4

    • peliostomi (Celonites)

    •  Celonites 11

    • peliostomi (Masarina)

    •  Masarina 19

    • peringueyi (Ceramius)

    •  Ceramius 16

    • perkinsi (Paragia)

    •  Paragia 7

    • perone (Quartinia)

    •  Quartinia 27

    • persephone (Quartinia)

    •  Quartinia 27

    • persicus (Celonites)

    •  Celonites 11

    • phaceliae (Pseudomasaris)

    •  Pseudomasaris 22

    • philomela (Quartinioides)

    •  Quartinia 27

    • phleomis]![ (Celonites)

    •  Celonites phlomis 11

    • phlomis (Celonites)

    •  Celonites 11

    • phoebe (Quartinioides)

    •  Quartinia 27

    • pictifrons (Paragia)

    •  Metaparagia 6

    • pictus (Celonites)

    •  Celonites 11

    • pictus rufiventris (Celonites)

    •  Celonites pictus ssp. 12

    • Plesiozethus

    •  Paramasaris 4

    • pluto (Quartinia)

    •  Quartinia 27

    • poecila (Quartinia)

    •  Quartinia 27

    • popovi (Quartinia)

    •  Quartinia 27

    • praedator (Paragia)

    •  Paragia odyneroides 7

    • Priscomasaris 5

    • Priscomasarina 5

    • promontorii (Celonites)

    •  Celonites 12

    • propinqua (Quartinia)

    •  Quartinia 27

    • propodealis (Paragia)

    •  Paragia 7

    • proserpina (Quartinia)

    •  Quartinia 27

    • Pseudomasaris 21

    • punctulata (Quartinia)

    •  Quartinia 27

    • purcelli (Celonites)

    •  Celonites 12

    • purcelli var. flavomaculatus (Celonites)

    •  Celonites purcelli 12

    • pusilla (Quartinia)

    •  Quartinia 28

    • puslilla[!] (Quartinia)

    •  Quartinia pusilla 28

    • Quartinia 22

    • Quartiniella

    •  Quartinia 22

    • Quartinioides

    •  Quartinia 22

    • rachiphorus[!] (Trimeria)

    •  Trimeria rhachiphora 30

    • reedi (Gayella)

    •  Gayella 4

    • rex (Ceramius)

    •  Ceramius 16

    • rhachiphorus (Odynerus)

    •  Trimeria rhachiphora 30

    • richardsi (Ceramius)

    •  Ceramius 16

    • richardsi (Paragayella)

    •  Paramasaris 5

    • Riekia

    •  Metaparagia 5

    • rohweri (Pseudomasaris)

    •  Pseudomasaris maculifrons 21

    • Rolandia

    •  Metaparagia 5

    • romandi (Erynnis)

    •  Masaris vespiformis 20

    • rothschildi (Celonites)

    •  Celonites 12

    • rotschildi[!] (Celonites)

    •  Celonites rothschildi 12

    • rudesculptus (Celonites)

    •  Celonites 12

    • rufomaculatus (Ceramius)

    •  Ceramius lichtensteinii 15

    • rugiceps (Celonites)

    •  Celonites 12

    • Saryara

    •  Masaris 20

    • Saryarinae

    •  Masarinae 4

    • saussurei (Masaris)

    •  Jugurtia 18

    •  Masaris carli 20

    • saussurei[!] (Paragia)

    •  Paragia tricolor 8

    • saussurii (Paragia)

    •  Paragia tricolor 8

    • savignyi (Celonites)

    •  ?Celonites fischeri 9

    • schulthessi (Ceramius)

    •  Ceramius cerceriformis 15

    • schulthessi (Paragia)

    •  Paragia 7

    • scutellimacula (Quartinia)

    •  Quartinia capensis 23

    • semenovi (Celonites)

    •  Celonites 12

    • senecionis (Quartinioides)

    •  Quartinia 28

    • separata (Quartinia)

    •  Quartinia 28

    • shestakovi (Quartinia)

    •  Quartinia 28

    • sicheliana (Gayella)

    •  Gayella eumenoides 4

    • signata (Quartinia)

    •  Quartinia 28

    • signata var. signatifrons (Quartinia)

    •  Quartinia signatifrons 28

    • simpsoni (Jugurtia)

    •  Jugurtia 18

    • smirnovi (Masaris)

    •  Masaris carli 20

    • smithii (Paragia)

    •  Paragia 7

    • sobrina (Paragia)

    •  Paragia 8

    • socius (Ceramius)

    •  Ceramius 16

    • soika[!] (Quartinia)

    •  Quartinia soikai 28

    • soikai (Quartinia)

    •  Quartinia 28

    • spinolae (Masaris)

    •  Jugurtia 18

    • spinolai[!] (Jugurtia)

    •  Jugurtia spinolae 19

    • spinosus (Celonites)

    •  Celonites 12

    • spiricornis (Ceramius)

    •  Ceramius auctus 14

    • spirolai[!] (Jugurtia)

    •  Jugurtia spinolae 19

    • striata (Quartiniella)

    •  Quartinia 28

    • strucki (Masarina)

    •  Masarina 20

    • syriaca (Quartinia)

    •  Quartinia 28

    • syriaca nitens (Quartinia)

    •  Quartinia syriaca ssp. 28

    • tarsata (Quartinioides)

    •  Quartinia 28

    • tenerifina (Quartinia)

    •  Quartinia 28

    • testaceopicta (?Masariella)

    •  Jugurtia simpsoni 18

    • texanus (Masaris)

    •  Pseudomasaris 22

    • thabaica[!] (Quartinia)

    •  Quartinia thebaica 29

    • thebaica (Quartinia)

    •  Quartinia 28

    • thebica[!] (Quartinia)

    •  Quartinia thebaica 28

    • tianshanicus (Masaris)

    •  Masaris 20

    • tigrina (Jugurtia)

    •  Jugurtia 19

    • titania (Quartinioides)

    •  Quartinia 29

    • toriger (Ceramius)

    •  Ceramius 16

    • Toryna

    •  Pseudomasaris 21

    • tricolor (Paragia)

    •  Paragia 8

    • tricolorata (Quartinia)

    •  Quartinia 29

    • Trimeria 29

    • tripolitana (Quartinia)

    •  Quartinia 29

    • tripolitana sinaitica (Quartinia)

    •  Quartinia tripolitana ssp. 29

    • tristiculus (Celonites)

    •  Celonites 12

    • tristiculus karatauicus (Celonites)

    •  Celonites tristiculus ssp. 12

    • tuareg (Quartinia)

    •  Quartinia 29

    • tuberculifer (Ceramius)

    •  Ceramius 16

    • tumidiscutellatus (Celonites)

    •  Celonites 12

    • turneri (?Masariella)

    •  Jugurtia 19

    • turneri (Celonites)

    •  Celonites 12

    • turneri (Quartiniella)

    •  Quartinia 29

    • turneri eburnea (Masariella)

    •  Jugurtia eburnea 18

    • tylecodoni (Masarina)

    •  Masarina 20

    • uzbeka (Quartinia)

    •  Quartinia 29

    • vagepunctata (Quartinia)

    •  Quartinia 29

    • variegata (Quartinia)

    •  ? Quartinia signata 28

    • varipennis (Celonites)

    •  Celonites 12

    • vechti (Ceramius)

    •  Ceramius 17

    • venusta (Paragia)

    •  Paragia 8

    • vespiformis (Ceramius)

    •  Ceramius cerceriformis 15

    • vespiformis (Cimbex)

    •  Celonites abbreviatus 8

    • vespiformis (Masaris)

    •  Masaris 20

    • vespiformis (Paragia)

    •  Paragia 8

    • vespiformis aegyptiacus (Masaris)

    •  Masaris aegyptiacus 20

    • vespiformis arabicus (Masaris)

    •  Masaris aegyptiacus ssp. 20

    • vespoides (Masaris)

    •  Pseudomasaris 22

    • vespoides robertsoni (Pseudomasaris)

    •  Pseudomasaris vespoides 22

    • wahlenbergiae (Celonites)

    •  Celonites 13

    • walkeri (Paragia)

    •  Paragia 8

    • waterstoni (Quartiniella)

    •  Quartinia 29

    • wheeleri (Celonites)

    •  Celonites 13

    • wheeleri (Pseudomasaris)

    •  Pseudomasaris 22

    • wheeleri var. immaculatus (Celonites)

    •  Celonites wheeleri 13

    • Willinkia

    •  Trimeria 29

    • yemenensis (Celonites)

    •  Celonites 13

    • yemenensis ethiopicus (Celonites)

    •  Celonites yemenensis ssp. 13

    • yemensis[!] (Celonites)

    •  Celonites 13

    • zarudnyi (Jugurtia)

    •  Jugurtia 19

    • zavattarii (Celonites)

    •  Celonites 13

    • Zethoides

    •  Paramasaris 4

    • zonalis (Masaris)

    •  Pseudomasaris 22

    • zonalis albopictus (Pseudomasaris)

    •  Pseudomasaris zonalis 22

    • zonalis basirufus (Pseudomasaris)

    •  Pseudomasaris basirufus 21

    • zonalis neomexicanus (Pseudomasaris)

    •  Pseudomasaris maculifrons 21

    JAMES M. CARPENTER "Checklist of Species of the Subfamily Masarinae (Hymenoptera: Vespidae)," American Museum Novitates 2001(3325), 1-40, (26 April 2001). https://doi.org/10.1206/0003-0082(2001)325<0001:COSOTS>2.0.CO;2
    Published: 26 April 2001
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